Glossary of Botanical Terms

This glossary of botanical terms is a list of terms related to botany and horticulture. These include flower terms, plant terms and words, botanical definitions and terminology.

What is botany? Botany is a branch of scientific study in which plants themselves are examined. If you are looking for general answers related to botanical and horticulture topics, education, job and carrier opportunities, check out our Botanical Frequently Asked Questions page.

-merous

A suffix preceded by an Arabic number or numeric prefix such as tri-, indicating how many of each part a flower possesses; 5-merous would be a flower with 5 sepals, 5 petals, etc., while trimerous would have three of each part.

A horizon

The uppermost layers of soils consisting of partly decomposed plant remains and relatively fresh leaves and other plant debris; the surface mineral layer, high in organic matter and dark in color; and the lighter colored layer where leaching of solutes and suspended materials occurs.

AARS

An acronym representing the name All-America Rose selections, Inc., an association of commercial rose growers that tests and approves new rose varieties for commercial use.

abaxial

Facing away from the axis.

abbreviated (syn. abridged)

Shortened, as when one part is shorter than another.

aberrant

Atypical or unusual; not normally occurring.

abiogenesis

Spontaneous generation of life from nonliving material.

abiotic

Nonliving, as opposed to biological.

abiotic stress

Nonliving environmental factors such as frost, drought, excessive heat, high winds, etc., that can have harmful effects on plants.

abiotic transformation

Any process in which a chemical in the environment is altered by nonbiological mechanisms, e.g., by exposure to sunlight.

abortion

Imperfect development or nondevelopment of an organ.

abortive

Defective or barren. Describes especially but not exclusively reproductive structures.
above. 1. Refers to the adaxial (upper/outer) surface of a leaf, petal, sepal or scale. 2. Refers to the part of a stem, bulb, tuber, branch, or inflorescence that is the greatest distance from the attachment, following the stem/branch; if a branch arches with a flower at the tip drooping nearly to the ground, the flower is said to be above the highest point of the branch.

abrade

To scrape off.

abrupt

Suddenly narrowed or cut off.

abscisic acid (abr. ABA, syn. abscisin)

A plant hormone that promotes leaf.

absciss-layer

A belt of tissue preparing for separation, especially at the connection of a leaf to the stem.abscission, fruit drop, and seed dormancy, and has an inhibitory effect on cell elongation.

abscission (adj. abscissile)

The normal shedding of leaves, flowers or fruit from a plant at a special separation layer, or abscission zone.

abscission zone

The area of separation when a plant sheds a leaf, flower or fruit.

absent petiole

A leaf without a petiole; sessile.

absorb (n. absorption)

To suck up or take up, e.g., plant roots absorb water.

absorbents

Antacid herb.

absorption spectrum

A graph of the amount of light a substance absorbs, plotted as a function of energy, frequency, or wavelength.

acariasis

An infestation with or a disease caused by mites.

acarodomatia

Domatia which have adapted to provide shelter to beneficial mites.

acaulescent

A plant that is stemless or apparently so, or with the stem subterranean.

accelerators

Admixtures that decrease the setting time of concrete by increasing the rate of hydration.

accessory

Auxiliary, subsidiary; as the parts of a flower beyond the necessary male and female organs, such as petals and sepals.

accessory bud

Buds that are at or near the nodes but not in the axils of the leaves.

accessory fruit

A fruit, or group of fruits derived from a single flower, in which the conspicuous, fleshy portion develops from the receptacle and is shed with the true fruit(s) attached.

accessory organs

Parts of a flower that are not directly connected with male and female organs, e.g., petals and sepals, etc.

acclimate (n. acclimation)

To adapt to a new environment or a change in the environment.

acclimatization (syn. adaptation)

The physiological process by which an organism adapts to a new environment.

accrescent

Gaining in girth or length with age or following fertilization, or growing together, as the calyx of some plants after flowering, such as the ground cherry.

accumbent

A cotyledon has the edges lying against the radicle.

aceriform

Shaped like a maple leaf; deeply lobed.

acerose

Sharp, solid, needle-like.

achene

A small dry and hard 1-celled, 1-seeded indehiscent fruit.

achenecetum

A cluster of small seeds (achenes), as in buttercups, Ranunculus.

achenocarp

Any dry fruit that does not open by itself.

achlamydeous

Without a perianth, e.g., the flowers of a willow.

achlorophyllous

Lacking chlorophyll; appearing without green color.

acicular

Needle-shaped. If solid may be either round or grooved in cross-section.

aciculate

Describes a surface that is etched with fine lines, as if scratched by a needle. 2. Marked with what appear to be pin pricks, usually arranged at random.

acid rain

Rain which has turned acidic due to the presence of sulfur or nitrogen oxides in the atmosphere. Acid rain can harm and even kill plants and aquatic life.

acid soil (adj. acidic)

Soil with a pH level below 7 is considered acidic; also called sour.

acidifier

An additive that decreases the pH (and increases the acidity) of soil.

acidophile (adj. acidophilic)

A microorganism that can, or must, live in an acidic environment. See also: alkaliphile.

acidulous

Slightly acidic.

acinaceous

Covered or filled with kernels.

acinaciform

Shaped like a half moon.

acinose

Resembling a bunch of grapes.

acondylose

Stems which lack joints or nodes.

acorn

The hard-shelled, one-seeded fruit of the oaks (Quercus) which consists of a nut which is partly enclosed by the cupule, or cap.

acotyledonous

Having no seed leaves, or cotyledons.

acre

Land measurement of 43,560 square feet, roughly the size of a football field.

acrid

Harsh and bitter in taste.

acrocarpous

In mosses, having the sporophyte terminal on a stem or ordinary branch.

acrocidal capsule

A dry fruit which dehisces by slits at the terminal.

acrodome

With the main veins ending at the leaf tip.

acrodromous

Describes leaves with two or more primary or strongly developed secondary veins running in convergent arches towards the apex. Arches not recurved at base. See also: brochidodromous, eucamptodromous, semicraspedodromous.

acropetal

Developing in a succession from the base and towards the apex. See also: basipetal

acroscopic

Facing upwards, towards the apex of the axis on which it is borne. See also: basocopic.

acrostichoid

Having sporangia apparently scattered on the surface of the fertile lamina.

acrotonic

Describes flowering seasonal shoots which produce leaves below the inflorescence. See also: basitonic.

actinomorphic

Refers to flowers that can be divided into symmetrical halves along any diameter. See also: zygomorphic.

actinostelic

Refers to a vascular strand where ribs radiate outward.

active transport

The movement of a chemical substance by the expenditure of energy through a gradient (as across a cell membrane) in concentration or electrical potential and opposite to the direction of normal diffusion.

actual

As in actual nitrogen; calculating the amount of the mineral present, e.g., calculating a 25-pound bag of fertilizer containing 22 percent of nitrogen as the equation 25 pounds X .22 = 5.5 pounds of actual nitrogen.

aculeate

Prickly; beset with prickles.

aculeiform

Shaped like a prickle.

aculeolate

Beset with diminutive prickles.

acumen

The gradually tapering narrow point of an acuminate leaf.

acuminate (n. acumination)

The shape of a tip or base of a leaf or perianth segment where the part tapers gradually and often in a concave manner.

acutangular

Refers to a stem that is sharply angled.

acute

Terminating with a sharp or well defined angle.

acyclic

Arranged in spirals as opposed to whorls.

adaptability

Proficiency of an organism to make changes improving survivability and reproduction in its habitat.
adaptation (alt. adaption)The ways an organism becomes better fitted to survive and reproduce in a particular environment. 2. A genetically changing characteristic that raises an organism’s ability to survive.

adaptedness

The genetic characteristics by which an organism is suited to its environment.

adaptive capacity

The genetically set range or flexibility of reactions of an organism enabling it to respond in different ways to differing conditions.

adaptive radiation

The evolution of new species or subspecies to fill unoccupied ecological niches.

adaptive selection

The evolution of comparable forms in separate but ecologically similar areas.

adaxial

Facing toward the axis, as the upper surface of a leaf.

additive

Material added to a substance, such as fertilizer, to better enable it to perform the desired function.

adenosine diphosphate (abr. ADP)

An ester of adenosine that is reversibly converted to ATP for the storing of energy by the addition of a high-energy phosphate group.

adenosine triphosphate (abr. ATP)

A phosphorylated nucleoside that supplies energy for many biochemical cellular processes by undergoing enzymatic hydrolysis especially to ADP. An energy-rich phosphorous compound that is important in the transfer of energy in organisms.

adherent


Joined, but not united.
adiabatic


Refers to an event in which heat is neither gained nor lost.
adjacent
Next to each other, but without touching or overlapping.
adjustment
Functional, never structural, changes by which an organism becomes better suited to its environment. See also: adaptation.
admixture
A material other than water, aggregates, or cement that is used as an ingredient of concrete or mortar to control setting and early hardening, workability, or to provide additional cementing properties.
adnate
United; fused, as the inferior ovary with the calyx-tube. Adnate anther: one attached for its wholelength to the inner or outer face of the filament.
adobe
A fine calcareous clay or silt, often used for construction purposes.
adpressed
Pressed flat against another organ.
adsorption
The attachment of molecules or ions to outer surfaces or interfaces.
advance growth (syn. young growth)
Seedlings and saplings appearing under the canopy ready to fill in forest openings made available by the death of a mature tree from logging, lightning kills, etc.
adventitious
Describes an organ growing where it is not normally expected, e.g., roots growing from a stem.
adventitious root (syn. stem root)
A root that arises from a stem, rather than from the primary root.
adventitious species
An alien or exotic species; an invasive species.
adventive
Describes an exotic species recently introduced to an area or imperfectly naturalized.
adventive taxa
Organisms that were not native to an area, and have now naturalized.
adze (alt. adz)
A tool for shaping wood having a curved blade arched downwards and at a right angle to the handle.
aelophilous
Refers to plants which spread seeds or pollen by wind.
aeolian (alt. eolian)
Refers to the wind, or to soil which has been moved by wind.
aerate
To supply or impregnate with air.
aerating root
A root structure that rises above ground, usually above water, to allow the plant to absorb air.
aeration
The processes by which air and other gases in a medium are exchanged or refreshed.
aerenchyma
Tissue with thin walled cells separated by large, gas-filled spaces that facilitates gaseous exchange and maintains buoyancy; typical of aquatic plants.
aerial
Refers to any plant part that is above water in aquatic plants; less often used with terrestrial plant parts found above ground.
aerial photograph
A map-like picture taken of the ground from high in the air, showing roads, fields, and other man-made objects as well as natural features such as rivers.
aerial roots

Roots borne wholly above ground, as the attachments of vine forms of Toxicodendron radicans which penetrate tree bark. 2. Rooting shoots of epiphytes.
aerobic (n. aerobe)
Living, active, or occurring only in the presence of oxygen.
aerobiosis
All life that grows in the presence of free oxygen.
aeroplankton
Microorganisms which float in the air, as plankton floats in water.
aerotaxis
An involuntary response of a living thing to a gas, such as a plant curving toward a more concentrated source of carbon dioxide.
aerotropism
An organism’s growth determined by the presence of oxygen.
aestidurilignosa
Woodland with mixed evergreen and deciduous hardwoods.
aestival (alt. estival)
Appearing during the summer season. See also: aspection.
aestivation

Estivation. 2. The arrangement of the perianth or their lobes in an unexpanded flower bud. See also: vernation.
affinity
That relationship between organisms which shows they share a common origin; used occasionally to show the similarity among communities.
affixed
Fastened upon.
afforestation
The process of starting a new forest in an area where none existed; to do the same to replace a previous growth is reforestation.
afoliate
Without leaves.
after-ripening

Metabolic changes that must take place in a seed to overcome dormancy. 2. The dormancy period following seed formation, necessary for embryo changes that insure germination.
aftermath

A second growth crop, also called a rowen. 2. The outcome, especially of a disaster, as in the aftermath of a forest fire.
agamic
Asexual.
agamogenesis
Asexual reproduction, as by budding, cell division, or parthenogenesis.
agamospecies (alt. agameon, syn. binom)
A group of individuals in which reproduction is almost exclusively by asexual means.
agamospermy
Apogamy where sexual union is not completed, yet the embryo is produced from the inside layer of the female gametophyte.
agamosporous
Having a life cycle in which chromosome segregation and recombination does not take place.
agamous
Without stamens or pistils; neuter, sterile.
agar
A gelatinous substance produced by red algae, often used as a culture medium.
agaric

The dried fruiting body of a fungus formerly used in medicine. 2. Any of a family (Agaricaceae) of fungi with the sporophore usually resembling an umbrella and with numerous gills on the underside of the cap.
age and area
The hypothesis that the larger the area covered by a species, the older that species is.
age class (syn. even aged)
Refers to a stand in which all the trees or other perennials started growth in the same regeneration period. See also: monoculture.
age distribution
The classification of groups within a community according to age or certain periods such as prereproductive, reproductive, postreproductive.
ageotropic (alt. apogeotropic)
Said of parts that would be expected to grow as gravity pulls, but instead grow upward, such as the knee roots of cypress, Taxodium spp./I>.
agglomerate
Together in a head, as the florets of red clover, Trifolium pratense; clustered.
agglutinate
Stuck together, as in pollen masses of orchids.
aggradation
The gradual filling of depressions in the earth’s surface by soil deposition, such as the deposition at a lake’s bottom; or the opposite, the wearing away of mountain tops, reaching toward a level surface.
aggregate

Clustered. In mosses, applied to two or more sporophytes from one perichaetium. 2. Composed of mineral crystals of one or more kinds or of rock fragments. 3. A material such as sand or gravel mixed with a binder, such as cement, to produce mortars and concrete. 4. A group of closely-related species.
aggregate flowers
Crowded into dense clusters or tufts, as in Scabiosa atropurpurea.
aggregate fruit
Describes a fruit, such as a raspberry, which consists of a fused cluster of several fruits, each one formed from an individual ovary. See also: syncarp.
aggregate species
A group of species that are so closely related that they are regarded as a single species.
aggregation
The coming together of organisms into a group, such as seedlings growing near the base of a parent tree. See also: community.
aging
When said of a lake, refers to the enrichment of waters, rapid growth of aquatic plants, and sedimentation which accelerate the death of a lake.
agonistic behavior (alt. agonistic behaviour)
Aggressive or defensive actions, such as fleeing or fighting, brought on by the interaction between individuals usually of the same species.
agrarian zone
That portion of a jurisdiction that can be or is farmed.
agrestal
Growing wild; especially weed plants growing on farms.
agrestic
Rural; rustic.
agrobiology
The study of plant nutrition and crop production in relation to soil control.
agroforestry
Land management for the simultaneous production of crops and trees.
agrology (syn. edaphology)
The study of soils.
agronomy (adj. agronomic)
The theory and practice of soil management and field crop production.
agrophilous
Refers to organisms which grow best in cultivated fields or other manmade areas.
agrostology
The branch of systematic botany which encompasses grasses; graminae.
aianthous
Flowers all year; semperflorus.
aiphyllus
Evergreen.
air layering (alt. air-layering, syn. Chinese layering, syn. gootee, syn. marcotting)
A method of propagation where a cut is made in a woody stem and surrounded by damp peat moss held in place by a wrap. When roots form, the stem can be removed and planted.
air mining
Processing air to recover minerals in suspension or solution.
akene
A dry, one-seeded indehiscent fruit, with the pericarp fitting closely around the seed.
ala (pl. alae)
Wing; in ferns, the narrow membrane laterally bordering a stipe, rachis, or costa.
alar
Describes the cells at the basal angles of the leaf, commonly different from the cells of the main part of the leaf, being shorter and often nearly square, or inflated and hyaline, and often highly colored.
alate (syn. aliferous)
Winged.
albedo
The proportion of incident radiation, usually light, that is reflected by a body such as a cloud.
albescense (adj. albescent)
The act of turning white; whitish.
albidus
Whitish.
albino
A flower that lacks normal color; white.
albumen
Starchy and other nutritive material in a seed, stored as endosperm inside the embryo sac, or as perisperm in the surrounding nucellar cells; any deposit of nutritive material accompanying the embryo.
albuminous
Having albumen.
albuminous cell
A gymnosperm phloem parenchyma cell closely associated with an adjacent sieve cell.
aletophyte
A plant growing in a mesic habitat.
alga (pl. algae)
A general name for the single-celled plant plankton, seaweeds, and their freshwater allies.
algae bloom
Rapid growth and death of aquatic plants, especially during hot weather in highly nutritious water.
algal layer
A thin layer of green or blue-green algae lying just beneath the cortex of a lichen.
algin
A gelatinous substance produced by brown algae, used in food and pharmaceutical preparations.
algoid
Resembling algae.
algology
The study of algae.
alien
An exotic; an introduced plant which has naturalized.
aliferous
Winged.
aliquote
The constant of temperature required for the development of a certain stage in the life cycle of an organism. See also: temperature summation.
alkali
A soluble salt obtained from the ashes of plants and consisting largely of potassium or sodium carbonate, e.g., a hydroxide or carbonate of an alkali metal, having marked basic properties. 2. Alkali metal. 3. A soluble salt or a mixture of soluble salts present in some soils of arid regions in a quantity that is detrimental to agriculture.
alkali reserve
The total amount of dissolved substances predisposed to maintain the normal alkalinity of a body of water or an organism’s internal body fluids.
alkali sink
A land basin in which water evaporation produces high salt concentrations that may, or may not, support salt marsh vegetation.
alkaline (adj. alkalinity)

Of, relating to, containing, or having the properties of an alkali or alkali metal. 2 A base; having a pH of more than 7.
alkaline soil (alt. alkali soil, syn. basic soil)
Soil with a pH level above 7; also called sweet.
alkaliphile (alt. alkalophile, syn. basophile, adj. alkaliphilic)
An organism that prefers, or can tolerate, alkaline conditions, typically in the range of pH 8-11. See also: acidophile.
allagostemonous
A flower having stamens attached to both the petal and receptacle.
allantoid
Sausage-shaped.
allee
A walkway lined with trees or tall shrubs.
allele (alt. allelomorph, syn. dominant character)
Any of the alternative forms of a gene that may occur at a given locus, one provided by each parent and determining charateristics such as eye color.
allelopathy (adj. allelopathic)
The inhibition of growth of one plant species by another due to the release of chemical substances.
allergen (adj. allergenic)
A substance which induces allergic symptoms like rash, inflammation, etc.
allergy
Sensitivity resulting in reactions such as rash, inflammation, etc., when exposed to an allergen, e.g., pollen, strawberries, etc.
alley
A garden or park walk bordered by trees, shrubs, or flower beds.
alliaceous
Having the smell or taste of garlic.
alliance
A group of plant associations classed together on the basis of similarities in floristic and sociological characters.
allochoric
Refers to a species occurring in two or more similar communities in the same region.
allochthonous
Originating from outside a system, such as the leaves of terrestrial plants that fall into a stream. See also: autochthonous.
allogamy
Reproducing by cross-fertilization.
allogenic succession
The replacement of one kind of community with another because of a change in the environment which was not produced by the plants themselves, e.g., a decrease in soil moisture.
allohexaploid
Having sex genomes with one or more sets derived from a species different from the other sets.
allopatric
Refers to two or more species having nonoverlapping ranges of distribution. See also: sympatric.
allopelagic
Refers to any organism occurring at any depth of the ocean.
allopolyploid
A polyploid individual or strain having a chromosome set composed of two or more chromosome sets derived more or less complete from different species.
allotetraploid
Having four genomes with two sets (rarely one) coming from a different species than the others.
allotrophic
Refers to lakes or ponds which receive organic materials from the surrounding land by washing.
alluvial
Describes something related to, or caused by, a river or stream, such as alluvial deposits.
alluvial fan
A delta at the mouth of a stream or river formed by the depositing of transported materials when the flow lessens.
alluvial soil
Soil deposits at the mouth of a stream or river, characterized by little or no modification of the original transported material by soil-forming processes.
alluvion
Detritus and sediments deposited by streams or the action of waves and currents in larger bodies of water; this contributes to the gradual addition of land and loss of lake or wetland.
alluvium
Sediments which were transported by a stream or river, then deposited when the stream flow lessened. See also: alluvion.
alm
A high mountain meadow, either alpine or subalpine.
almanac
Calendar to which astronomical data and various statistics are added.
alpage
An upland pasture consisting of natural plants which are used for grazing animals at the height of summer.
alpine

A plant native to a mountainous region. 2. Refers to the area of a mountain above the tree line, or to organisms which grow there.
alteratives
Herbal medicines which can work a gradual change in the system.
alternate
Describes leaves that are not opposite to each other on the axis, but arranged singly at different heights.
alternation of generations
The complete life cycle of plants with very different sexual and asexual phases, such as the successionof haploid gametophytes and diploid sporophytes found in ferns, Pteridophyta.
alternes
Two or more types of communities alternating with each other in a more or less restricted area.
alternitepalous
Describes floral parts, inserted alternately with the tepals.
altherbosa
Communities with tall herbaceous plants, especially in denuded forest areas.
altimeter
An instrument for determining altitude.
alulate
Having a very narrow wing.
alvar
A Swedish term for a habitat type consisting of dwarf shrubs.
alveolate
Honeycombed; having angular depressions separated by thin partitions.
alveolation
A honeycombed condition.
amanthophilous (syn. ammocolous, alt. ammophilous)
Refers to organisms which live in sand.
amendment
Alteration or change, especially for the better, as when adding a soil amendment.
amensalism
The state or interaction in which one organism is reduced while another is not influenced. See also: commensalism.
ament
A catkin, or scaly spike.
amentaceous
Refers to plants which produce catkins.
amentula
Applied to the special antheridia-bearing branches of Sphagnum.
American Rose Society (abr. ARS)
An educational, nonprofit organization dedicated exclusively to the cultivation and enjoyment of roses.
amethystine
Violet-colored.
ametoecious
Refers to a parasite which is restricted to a single host, i.e., species specific.
amino acid
One of the organic, nitrogen-containing units of which proteins are composed.
amitosis
Direct division of the nucleus of a cell without mitosis.
ammate
A compound of ammonium sulfate, often used as an herbicide.
ammonification
The formation of ammonia compounds from organic materials containing nitrogen.
amnion (alt. amniotic fluid, alt. amniotic waters)
The liquid surrounding an embryo while in uterus or in an egg.
amniote
Animals whose embryos develop within a fluid-filled sac, e.g., some reptiles, birds and mammals.
amorphous
Shapeless, a form not regular or predictable.
amphibious
Capable of living on land or in the water.
amphicarpic
Producing two different types of fruits; sometimes said of plants which bear more than one crop each year.
amphicarpogenous
Producing fruit above ground which then becomes buried, e.g., peanuts, Arachis hypogaea.
amphichromatism
Having flowers of different colors appearing in different seasons.
amphicryptophyte
A marsh plant with some underwater vegetative parts.
amphidiploid (alt. amphiploid)
A tetraploid having two sets of genomes coming from different parent stock.
amphigeal
Producing two types of flowers, one coming up directly from the root, the other on upper stems.
amphigean
Native of both Old and New Worlds.
amphigenesis
The joining of gametes to form a zygote.
amphigenetic
Refers to an organism or a part of the growth cycle that produces zygotes.
amphimixis
Sexual reproduction; the joining of parental characters.
amphiphyte
A plant growing in the edge of wetland and water showing amphibious characteristics.
amphiploid (alt. amphidiploid)
A variation on polyploid in which there are two sets of chromosomes, each set coming from a different species.
amphirhinal
Refers to an organism with two noses.
amphitheater (alt. theatre, syn. theater)
A hillside or depression with tiers or terraces cut into it, resembling a classical outdoor theater and usually used for that purpose.
amphithecium
The outer layers of cells of the sporangium.
amphitropous
Half inverted and straight, with the hilum lateral; an ovule that is curved back along its funiculus so that the base and micropyle are close together. Describes an ovule or a seed.
amphoteric
Refers to the ability of a substance to react either as an acid or a base.
amplectant
An organism or organ with the ability to clasp or twine, like a tendril.
amplexicaul
Clasping the stem.
ampliation
The process of enlarging or amplifying.
ampule (alt. ampulla, adj. ampulliform, adj. ampullaceous)
A hollow flasklike organ shaped like a bladder or squat round bottle; e.g., the traps and floats such as those found on the leaves of Nepenthaceae or Utricularia.
anabiont
A plant that produces flowers and fruits many times; a perennial.
anabiosis
Revival of an organism after apparent death, like the resurrection plant, Anastatica.
anabolism
The process of building up protoplasm from simple substances.
anadromous
Having the first lobe or segment of a pinna or frond arising toward the apex/tip.
anaerobe
An organism which can survive in the absence of oxygen.
anaerobic
Describes a process that occurs with little or no oxygen present.
anaerobic respiration
Gaseous exchange in which the hydrogen removed from the glucose during glycolysis is combined with an organic ion instead of oxygen.
anaerobiosis
The existence of life without oxygen.
anaerophytobionts
Soil flora which exist without oxygen.
analgesic
An herbal medicine which can relieve pain.
analogous
Of similar function, but of different evolutionary descent. See also: homologous.
anamniote
An animal lacking embryonic membranes or amnion.
anandrous
Refers to flowers that have no stamens.
anastomose (v. anastomosing, adj. anastomosed)
Netted, as are veins in a leaf.
anastomosing
Connecting by cross-veins and forming a network.
anastomosis
The connection of various parts to form a network, as in leaf veins.
anatomy
The study of the structure of an organism and the relationship between its parts.
anatropous
An ovule that is inverted and straight, with the micropyle next to the hilum and the radicleconsequently inferior.
anauxotelic
Describes inflorescences, parts of inflorescences or axes that do not end in a flower, and in which growth does not continue beyond the flowering region. See also: auxotelic.
anchor root
A large root serving mainly to hold a plant in place in the soil.
anchor-ice
Frozen water that forms at the bottom of a stream.
ancipital (adj. ancipitous)
With both edges sharp, as some flattened stems.
androconia
Modified scales on the wings of butterflies, e.g., Lepidoptera, that produce a sexually attractive odor. See also: pheromones.
androdioecious
A plant breeding type in which a species, to produce seeds, must have a male plant with flowers having only stamens and a bisexual plant with flowers having both stamens and pistils. See also: dioecious.
androecium
Refers collectively to the stamens of one particular flower.
androgynophore
A stalk bearing both the androecium and gynoecium of a flower above the perianth.
androgynous

Of an inflorescence composed of both staminate and pistillate flowers. 2. With antheridia and archegonia in the same cluster of leaves, i.e., either synoicous or paroicous.
andromonoecious
Refers to species that have bisexual and male flowers on the same plant.
androphile
A plant that grows most successfully around humankind.
androphore
A stalk bearing the androecium.
androsporangium
The receptacle in which androspores are formed.
androspore
The minute reproductive body, which gives rise to the (often exceedingly obscure) male plantlet in the sexual generation.
anemochore (alt. anemochory)
Dispersal of organisms, such as seeds, by wind. See also: diaspore, disseminule.
anemogram
The graphic record of wind velocity made by an anemograph.
anemograph
An automatic instrument for graphic recording of wind sp.ed.
anemometer
An instrument that measures wind sp.ed, but does not necessarily record it. See also: anemograph.
anemophious (alt. anemophilous)
Refers to plants which are pollinated almost exclusively by wind. See also: anemochory, entomophilous.
anemoplankton
Microorganisms transported by wind.
anemotaxis
The reaction to wind experienced by a free-moving organism.
anemotropism
Movement or growth of cells or organisms in response to wind.
aneroid barometer
An instrument that precisely measures atmospheric pressure to allow calculation of altitude.
aneuploid
Refers to the presence of an irregular number of chromosomes, higher or lower than multiples of the haploid number. See also: euploid.
anfractuose (alt. anfractuous)

Tightly twisted together. 2. Closely sinuous.
angiosperm
A flowering plant with ovules contained inside the ovary.
angiospermous
Having the seeds borne within a pericarp.
angle of repose (syn. critical slope)
The maximum degree of gradient on which soil or loose rock remain stable.
angled
Having evident ridges.
angular (alt. angulate)
Angled. Refers to an organ that shows a specific number of angles, e.g., mints, Labiatae, which have stems that are 4-angled, and are square in the cross section.
angular cells
See also: alar.
angustifolia
Narrow leaves.
angustiseptal
Having a fruit with narrow partitions, as the silicle of Cruciferae.
angustiseptate
Having narrow partitions. See also: latiseptate.
anhydrous
Refers to a substance which contains no water, like anhydrous ammonia.
animal unit
A standardized measure to understand the needs of different kinds of livestock in relation to forageresources. A mature cow of about 1000 pounds (455 kg.) is the standard unit, and the feed needed is equal to one horse, one mule, five sheep, five swine, five goats, or eight geese.
animal unit month
A measure of forage needed to maintain one animal unit for 30 days.
anion
A negatively-charged ion, such as chlorine. See also: cation.
aniso-
A prefix meaning unequal or dissimilar.
anisophyllous
Refers to paired leaves which are different in size or shape, common in trailing stemmed gesneriads.
anisophylly
The presence of two kinds of leaves on one plant as in Selaginella and some cedars, Juniperus. See also: dimorphism.
annotinal
Appearing yearly.
annual
A plant whose life cycle is of only one year’s duration.
annual production
The amount of yield each year by an organism or group.
annual ring
The layer of wood produced by a single year’s growth of a woody plant.
annual seasons
The major climatic periods of each year: vernal, estival, autumnal, and hibernal.
annual succession
The routine occurrence of plants and animals in an area during each year, such as spring bulbs being replaced by annual flowering plants, and subsequently replaced by autumn perennial flowering plants.
annual turnover
The total accumulation of living organisms produced in one year for a certain area. See also: biomass.
annuation
The highs and lows from year to year in abundance or performance of organisms caused by differences in environmental conditions such as temperature and moisture.
annular
In the form of a ring.
annular corona
Raised fleshy tissue, usually in a ring, on the corolla around the base of the staminal column but not closely adnate to it.
annulated
Made up of rings.
annulus

A ring of cells of the leptosporangium (the sporangium of higher ferns), which causes the sporangium to open and to discharge its spores.
anodynes
Herbal medicines that can counteract pain but are milder than analgesics.
anomalicidal capsule
A dry fruit which dehisces irregularly; a rupturing capsule.
anoxia
Condition resulting from an extreme lack of oxygen, usually resulting in permanent damage.
anoxic
Greatly deficient in oxygen.
antagonism
The depressive effect of one organism upon another, such as certain grasses like timothy hay on the production of alfalfa hay.
antecedent moisture
The degree of liquid which has soaked into the soil at the start of the runoff period.
antemarginal
Inside of, or not extending quite to the margin.
anterior
Away from the stem, midrib, pistil or other organ. On a lipped flower, the bottom lip is anterior, the top lip is posterior. Sometimes the anterior lip is less correctly called inferior or exterior.
antero-lateral
In front and on the sides.
anthelmintics
Herbal medicines that can destroy or expel intestinal worms.
anther
The polliniferous part of a stamen.
anther sac
A pocket-shaped unit containing pollen. In many plants, the anther has two lobes, each with two pollen sacs.
antheridium (pl. antheridia)
In cryptogams, the organ corresponding to an anther that produces male gametes; the male organ on the prothallium.
antheriferous
Anther-bearing; containing anthers.
antheroid
Similar to an anther.
antherozoid
Sperm, male gamete; one of the minute organs developed in an antheridium.
anthesis
The time of expansion of a flower.
anthocarp
A pseudocarp consisting of the true fruit and the base of the perianth.
anthocyanin (adj. anthocyanous)
A blue to purplish-red coloring agent, water soluble.
anthracnose
A fungus that forms grayish/whitish spots on leaves and stems.
anthropic (alt. anthropeic)
Refers to the influences of humankind in contrast to natural influences, such as the introduction of IPM pest control.
anthropochore (alt. anthropochory)
Dispersal of organisms, such as seeds, as a result of human activity. See also: anemochory.
anthropogenous
Refers to practices of humans, such as cultivation or monoculture.
anthropophilous
Refers to plants which grow near humans and their dwellings, such as dooryard violet, Viola odorata.
anti-
Against, opposed to.
anti-inflammatory
Refers to herbal or other medicines that can ease or neutralize swelling, heat, and pain.
antiallergenic
Refers to herbal medicines that can reduce or relieve allergic reactions.
antiaphrodisiac
Refers to herbal medicines that can reduce sexual desire.
antibiosis
The reaction, often death or sterilization, produced in organisms by an antibiotic.
antibiotic
Substance that slows the growth or multiplication of, or kills, a living organism; usually referring to bacteria.
anticline
A geological structure or arch formed by strata from opposite sides rising upward in a common line. See also: syncline.
anticyclone
An air mass with high atmospheric pressure surrounded by lower pressure areas, in which the circulation of the air is clockwise in the northern hemisphere and counterclockwise in the southern hemisphere.
antifungal
Refers to herbal medicines that can slow the growth or multiplication of fungi, or kill them.
antihistamine
Herbal or other medicines that can neutralize the effect or inhibit production of body histamines.
antimicrobial
Refers to herbal or other medicines that can slow the growth or multiplication of microorganisms, or kill them.
antipetalous (alt. antepetalous)
Opposite the petals.
antique rose
A rose which has been in cultivation since at least before 1900. Often refers specifically to an Old Garden Rose.
antiscorbutics
Refers to herbal medicines that provide Vitamin C.
antisepalous (alt. antesepalous)
Opposite the sepals.
antispasmodics
Refers to herbal medicines that can relieve cramping or spasms.
antitumor
Refers to herbal medicines that can prevent or be effective in removing tumors.
antitussive
Refers to herbal medicines that can prevent or ease coughing.
antiviral
Refers to herbal or other medicines that can slow the growth or multiplication of viruses, or kill them.
antrorse
Pointed forward and upward.
aperiodicity
Irregular occurrence of various phenomena, such as leaves dropping out of season (in summer rather than autumn) due to unexpected climatic changes, i.e., storms, droughts, etc. See also: periodicity.
apetalous
Having no petals.
apex
The top or tip of a structure.
aphids
Small, often wingless insects that suck sap.
aphotic
The deep zone of an ocean or lake receiving too little sunlight to permit photosynthesis.
aphotic zone
The deeper portions of bodies of water in which sunlight does not penetrate with enough intensity to cause changes in organisms. See also: disphotic zone, euphotic zone, photic zone.
aphototropism
The response of an organism in turning away from the source of light. See also: photophobic, photomanic.
aphrodisiac
Refers to herbal preparations that can stimulate sexual desire.
aphyllous
Without leaves.
aphytal zone
The deeper portion of a lake bottom that lacks plants, including the sublittoral zone and the profundal zone.
apical (adj. apically)

Borne at the tip of an organ, farthest from the point of attachment, e.g., a bud which terminates a stem. 2. Describes the cells composing the apex of the leaf. They are often broader and shorter than the cells of the middle of the leaf.
apical dominance
The ability of the apical meristem to produce hormones to prevent side shoots or buds from developing while it is growing.
apiculate
Having an apicule.
apicule
A short sharp point, not rigid, found at the tip of a leaf, bract, or petal.
apiculum (adj. apiculate)
A short, pointed, flexible tip.
apo-
A prefix denoting away from or separate.
apocarpous
With carpels separate rather than united. See also: syncarpous.
apogamous
Developed without fertilization.
apogamy
The formation of a sporophyte from a gametophyte by asexual means such as budding, rather than by sexual fertilization.
apogeotropism
The response by an organism of turning away from the earth, e.g., plant stems growing upward.
apomict (adj. apomictic)
A plant that reproduces by apomixis.
apomixis
Reproduction without fertilization or formation of gametes.
apomorphic
Describes a derived characteristic. See also: autoapomorphic, pleisiomorphic, synapomorphic.
apopetalous
With separate petals, not united to other petals; choripetalous.
apophysis
See hypophysis, the more correct term according to Braithwaite.
aposepalous
With separate petals, not united to other petals; chorisepalous.
aposporous
Having the condition of apospory.
apospory
The formation of a gametophyte from a sporophyte by asexual means without meiosis or sporeformation.
apothecium (pl. apothecia)
The open fungus fructification, as a cup or disc, lined with asci.
appendage
A structure attached to or arising from a larger structure.
appendiculate
Describes cilia with small transverse spurs attached at intervals along the margin. As these barssometimes extend inward instead of laterally, they are not always visible in a strictly dorsal view.
appendix
The long narrow development of the spadix in Araceae.
applanate
Pressed; flattened.
appressed
Lying close and flat against, as a bud against a twig.
approximate
Located close together and nearly touching.
apron
A layer of hard material like concrete or timber that protects soil from erosion, e.g., pavement at the spillway of a dam.
apterous
With no wings.
apud
The connector term used between names when two authors describe the same species in separate publications, e.g., Ceratozamia latifolia Miq. apud Tijdschr.
aquaculture

The raising of fish or other aquatic animals for their commercial value. 2. Hydroponic horticulture.
aquatic plants (alt. aquatics)
Those species whose seeds germinate in water or in bottom soil of bodies of water, usually with submersed or floating leaf types.
aqueduct
A conduit for water, whether canal, pipe, tunnel or a combination of these or others.
aquiculture
The use of artificial reefs and other structures to increase the production of seafood in fresh or salt water.
aquifer (alt. aquafer)
A natural holding tank of porous rock or soil locked between impermeable layers in which water may travel long distances.
aquiherbosa
Communities of herbaceous plants growing in ponds and marshes.
aquiprata
Communities of plants where the surface water is a necessary factor, as in wet meadows.
arable land
Land that may immediately be used for farming without clearing trees, draining, etc.
arachnoid (adj. arachnose)
A surface with many fine hairs that appears covered with cobwebs.
araneous
Cobwebby.
arbor (alt. arbour)

A lattice-covered structure with open sides, usually with vines, such as grapes trained across the top. 2. An orchard or a garden devoted to trees.
arboreal
Of or resembling a tree.
arboreous

Having many trees. 2. Tree-like.
arborescent
Tree-like in appearance and size.
arboretum (pl. arboreta)
A botanical center devoted to the study and exhibition of trees.
arboriculture
The cultivation of trees.
arcade
A series of arches with columns or piers; in gardens, usually having trees forming the arched ceiling.
archaeophyte
A plant that existed in prehistoric times.
archegone
The egg cell produced in the archegonium.
archegonium (pl. archegonia, adj. archegonial, adj. archegoniate)
The flask-shaped female reproductive organs on the prothallium in the higher cryptogamscorresponding to a pistil in the flowering plants and containing the egg which becomes the sporophyte. See also: antherozoid.
archibenthic zone (adj. archibenthal)
The ocean layers between 200 feet and 3300 feet (65 and 1050 m.)l; the upper part of the abyssal zone.
arching
Curved gently outward and then downward; generally said of stems, large leaves, and floral clusters.
architectural
Describes plants that have very strong shapes and are used in landscapes for this reason.
arctic
Describes an extremely cold climate, particularly that of the polar regions.
arctic-alpine
Refers to areas which are mountainous, above the tree line, and north of the arctic circle.
arcuate

Moderately curved. 2. In mosses, refers to a capsule bent in a curve like a bow.
area
The total range in which a taxon or community may be found. See also: basal area, coverage.
areg
A sand desert.
arenaceous

Describes a sandy soil. 2. Growing in sandy soil.
arenicolous
Refers to organisms which live where there is sandy subsoil.
areography
The study that deals with the range of a species. See also: area.
areola (pl. areolae)
A space enclosed by anastomosing veinlets.
areolate
Having areolae; marked out into small spaces; reticulate.
areolation
The network formed by the outlines of the cells of a leaf.
areole (adj. areolate)

The space between veins of a leaf or a similar partition. 2. A small pit or cavity marked out upon a surface. 3. The area from which hairs, spines, flowers, or branches may arise in Cactaceae.
argillaceous
Like clay, growing in clay, or clay-colored.
arid (n. aridity)
Xeric, extremely dry.
aril
A fleshy appendage growing at or about the hilum of a seed.
arillate
Having an aril.
arista (pl. aristae)
A bristle-like appendage, like the awn of grasses.
aristate (syn. awned)

Describes a leaf which terminates in a stiff, bristle-like tip. 2. Awned; provided with stiffish bristle-shaped appendages.
aristulate
Having a small awn.
armature

The basic framework of an object or organism. 2. A protective covering or other kind of defense, e.g., thorns.
aromatic
Having a spicy odor, at least when crushed.
aromatics
Refers to herbal medicines that have a pleasing odor and pungent taste.
arrangement
A nontechnical term that refers to the way things are put together, e.g., an inflorescence may be described by the arrangement of the flowers, or leaves can be arranged opposite or alternate.
arrhythmic
Refers to activity which is not dependent upon light or dark. See also: diurnal, nocturnal.
arroyo
A watercourse, gully, or channel carved by water, often dry.
ARS
An acronym representing the name the American Rose Society, a nonprofit, educational organization dedicated to the cultivation and enjoyment of roses.
arthropod
Insect, crab, spider, centipede, or other animal from the phylum Arthropoda.
article
A segment of a jointed stem.
articulate (syn. articulated)

Having a swollen area, often discolored, at the node. 2. With a joint, often on a zigzag stem.
artifact

A structure or appearance in a tissue due to death or the use of a reagent, and not present during life. 2. A product of human workmanship found on archeological digs.
artificial selection
The intentional human manipulation within a population to produce a desired evolutionary response. See also: natural selection.
artificial stocking
The introduction of animals from another area, such as providing streams with new species of fish or bringing gamebirds into an area where they are scarce.
arundinaceous
Reed-like.
ascending
Rising somewhat obliquely, or curving upward. Ascending ovule: one that is attached above the base of the ovary and is directed upward.
ascogonium
The special hypha from which the fertile hyphae develop to produce asci.
ascomycete
A fungus producing asci.
ascus (pl. asci)
The reproductive cell (meiotangium) of a fungus containing usually eight ascospores.
asepalous
Without sepals.
aseptate
With no partitions or divisions.
asexual
Lacking sexual characteristics as in a sterile ray floret; or when referring to reproduction, occurring without the fusion of egg and sperm.
asexual propagation
The propagation of plants through means other than fertilization, including layering, cuttings, tissue cultures, and the division of clumps.
asexual reproduction
Not involving or requiring fertilization and meiosis.
ashlar

Dressed, finely-jointed stonework, usually having a polished surface and sometimes consisting merely of a facing for a wall of rubble or brick. 2. The stones used in this stonework.
aspect

The appearance of vegetation during one of the seasons of the year, e.g., the vernal aspect. 2. The direction toward which a slope is facing, e.g., the southern aspect.
aspection
The change in the appearance of vegetation and its visible attributes during the succession of seasons of the year, as budding in prevernal, flowers in vernal, green fruit in estival, ripe fruit in serotinal, and bare branches in hibernal.
asperous
With a rough surface.
asperulate
Somewhat rough to the touch.
asperulous
Refers to a surface with short, hard projections.
assimilation
Cellular conversion of raw materials into structures useful to a plant, such as cell walls and protoplasm.
association complex
A group of associations which occurs in a defined area.
association fragment
A stand of plants that requires other characteristics to meet community standards.
association segregate
A climax community which has changed from the original community, like a mature beech-maple association appearing from a mixed deciduous forest.
association table (syn. synthesis table, syn. stand table)
A list of species that occurs in several stands of an association or community, including data on characteristics such as abundance, cover, vitality, etc.
associes
A temporary community in developmental stage.
assortative breeding (syn. assortative mating)
The pairing of male and female organisms in a manner that involves more than chance, so that the mating of similar parents is encouraged.
assurgent
Ascending.
asterean
Belonging to or similar to plants of the aster family, Compositae.
astomous
In mosses, refers to a capsule without a mouth. Used of capsules which have no regularly dehiscent lid or operculum.
astragaloid
Cubical.
astringent
An herbal preparation that shrinks and firms tissue, particularly the skin.
astylous
Lacking a style.
asymmetrical
Having a different shape on each side of a central axis.
asymptotic population
The maximum size possible to a population under present conditions, no matter how long reproduction is allowed to continue.
atavistic
Reverting to a form found in ancient ancestors.
atmometer
Any instrument for measuring evaporation like a porous sphere or open pan of water.
atoll
A circular coral island or islands surrounding a body of water in the ocean. See also: lagoon.
atrocastaneous
Very dark chestnut in color.
atrophy (adj. atrophied, alt. atrophic)

A wasting away from lack of nutrition or use. 2. Arrested development of a part or organ inconsequential to the normal development of a plant or animal.
atropurpureus
Colored deep purple, almost black.
attenuate
A long gradual taper, as at the base or tip of a leaf or flower parts, drawn out into a long point. See also: acute.
atypical
Not normal.
auctoris (alt. auctorum, abr. auct., abr. auctt.)
Used in taxonomy when an author has applied a wrong name, usually beginning with “non,” to contrast it to the true type with the correct author, e.g., Betula platyphylla auct. Non Sukachev, which is in fact a different plant, Betula mandschurica.
auger
A T-shaped tool, like a large corkscrew, used for drilling holes in the soil.
auricle
An ear-shaped appendage.
auriculate
Shaped like an ear.
auriculiform
Having the shape of an ear.
aurieles
Small lobes at the basal angles of the leaf, usually consisting of cells differing in size, shape, or both from those of the main part of the leaf. Properly used only when there is an outward curve in the outline of the leaf at the base, but often used loosely to denote the basal angles of widely decurrent leaves.
auroral
Refers to morning, the crepuscular period, or dawn.
autapomorphic
Describes a derived characteristic unique to a given taxon or monophyletic group. See also: apomorphic, synapomorphic.
autecology
The study of the individual in relation to environmental conditions, or sometimes, members of a speciesstudied collectively in the same way.
author

The botanist who discovered and named the new taxon. 2. The name or initial(s) following the taxon name, designating that botanist.
autochore
A species in which some action of the parent plant is the chief force for dissemination, e.g., the mechanical projection of seeds in jewelweed, Impatiens.
autochthonous

Describes the original or earliest known inhabitants of a region. 2. Originating in the place where found; indigenous. 3. Originating from within a system, such as organic matter in a stream resulting from photosynthesis by aquatic plants. See also: allochthonous.
autoecious (alt. autocious)

Refers to parasites which pass all stages of their life cycle on or within the same host, like certain rustfungus. 2. Having male and female organs on the same plant. See also: heteroecious.
autogamous
Self-fertilizing.
autogamy
Where male parts can fertilize female parts of the same flower through self-fertilzation or hermaphroditism; inbreeding.
autogenic succession
Vegetational progression in which each stage modifies the habitat in such a way that it is replaced by another stage, e.g., pond shoreline herbaceous plants being replaced by shrubs.
autolysis
The digestion of an organism or parts of it by its own enzymes.
autonomic
Refers to processes or activities that are spontaneous, arising from internal causes. See also: autonomous.
autonomous

Autonomic. 2. Refers to plants, especially those with chlorophyll, that are capable of turning inorganic materials to organic ones for their nutrition or other use. See also: autotrophic, photosynthesis.
autonym
When an author names a new subspecies or variety, the species is given the same new rank, based on the original type of the species and duplicating the epithet, e.g., when Pinus nigra ssp. larico was designated, Pinus nigra ssp. nigra came into being.
autopolyploid
An organism or cell which contains three or more sets of homologous chromosomes that developed from the same species. See also: amphiploid, allopolyploid.
autotrophic (n. autophyte)

Refers to organisms that are capable of processing inorganic materials into organic ones by using energy from outside the organism such as sunshine on chlorophyll. 2. Refers to a body of water that is limited in its supply of organic material to what it is capable of producing within its own borders.
autumnal (syn. fall)
Refers to the season between summer and winter that, in the northern hemisphere, includes the months of September, October, and December. Astronomically speaking, it is the period extending from the September equinox to the December solstice.
auximone
A nonessential organic substance of unknown chemical composition, which produces a known reaction, such as growth of duckweed, Lemna, from an extract of dung.
auxin
One of several hormones occurring in plants which regulates certain aspects of growth. They can be produced synthetically and are sometimes used commercially as herbicides or to promote flowering or other types of plant growth.
auxospore
A cell that restores the original size to the diminishing products of cell division.
auxotelic
Describes an inflorescences, parts of inflorescences or axes that do not end in a flower, and in which growth continues beyond the flowering region, See also: anauxotelic.
available nutrient
The portion of nutrient substances, such as nitrates in the soil, that can be utilized by plants at rates and amounts required for growth.
available water
That portion of water held in soil that may be absorbed into the roots of the plant.
available water-holding capacity
The amount of water in soil that can be absorbed by plants, between the high amount at full saturation or field capacity, and the low amount at the permanent wilting capacity.
avalanche
A large mass of snow, ice, rocks, trees, etc., in swift descent down a mountain or over a precipice.
avalanche cone
All the materials left at the base of a slope following an avalanche.
avalanche wind
Moving air stirred up by an avalanche and often destructive over considerable distances.
avenue
A path or road lined with trees on both sides, usually all of the same type.
average distance
The mathematical statement of the distance between plants calculated by dividing the square root of an area by the density of each species within the area.
avitaminosis
Reduced health from deficiency of vitamins.
awl-shaped
Tapering upward from the base to a slender or rigid point.
awn
A bristle-shaped appendage, especially on grass seeds or grains.
axe
A waist-high tool with a vertical, sharpened, narrow wedge head for cutting wood.
axe-shaped
Dolabriform; describes three-dimensional shapes.
axial
Refers to the main line or trunk of the entire plant, the specific branch, the inflorescence, etc.
axil
The angle formed by a leaf or branch with the stem.
axile
Belonging to, or found in, the axil.
axillary
Situated in an axil.
axis
The main line of growth in a plant or organ, e.g., the stem, from which the other parts such as the leaves and flowers grow.
azonal
Refers to soil that has no well developed profile with horizons, such as glacial deposits, dune sand, alluvium.
B horizon
The soil layer between A and c horizons, in which materials from overlying horizons accumulate from precipitation or suspension.
baccate
Berry-like; pulpy throughout.
bacciform
Berry-shaped.
bacilliform (syn. baculiform)
Rod-shaped.
Bacillus thuringiensis (abr. Bt)
A bacterium which is lethal to some insects and is used as a biological control.
backbulb
A dormant water-storing thickened stem that looks like a bulb, found in Orchidaceae. It grows actively as a pseudobulb the first year, then becomes dormant when the next year’s pseudobulb takes over.
backcross (alt. back cross)

To cross a first-generation hybrid with one of the parental types. 2. The offspring of a cross between a hybrid and one of its parents.
backfill
Earth used to fill trenches or holes, often the same soil that was removed to make the hole.
backfire
A fire purposely set ahead of an advancing fire to destroy flammable materials, enabling workers to control the main fire.
backflash
Movement of a poison through natural root grafts, from trees that have been treated with poison, back to non-treated trees.
backshore
That part of the beach from the first crest toward the land.
bactericide (alt. bactericidal)
Something, such as an herbal preparation, capable of destroying bacteria.
bacteriophage
A virus which destroys bacteria.
bacteriorhizae
Nodules on the roots of most legumes and a few other plants, which contain bacteria that uses atmospheric nitrogen to synthesize organic compounds.
baculiform (syn. bacilliform)
Rod-shaped.
badlands
Regions of eroded land on which most of the surface is covered with ridges, gullies, and deep channels, having sparse vegetation.
Baermann funnel
A modification of the Berlese funnel, used to force nematodes out of soil or debris by filling the funnel with warm water, driving the nematodes into a vessel below.
baffle-pier
A blockage placed in the path of high-velocity water, like a pier on the apron of an overflow dam.
baffles
Vanes, guides, or other devices inside a conduit to check eddy currents below them, and provide more uniform distribution of sp.ed.
bagasse
Residue of sugarcane after the juice has been extracted.
baguio
A term used in the Philippine Islands for a tropical cyclone or typhoon.
bajada
A term used in the southwestern united States to describe outwash slopes with long straight profiles.
balance of nature (alt. ecological balance, alt. biotic balance, syn. dynamic equilibrium, syn. biotic equilibrium)
A term for an ideal condition in which the interrelationships of organisms to one another and their environment appear harmonious, like a climax forest. In reality, the balance is continually upset by natural events.
balanoid
Refers to barnacles.
bald
A treeless region in forest vegetation; perhaps an area of grasses or shrubs in southern Appalachia, or a mountaintop.
balled and burlapped
Taken out of the ground with a ball of soil around the roots and then wrapped in burlap for support.
balling
A condition where outer petals stick together and fail to open, often occurring in damp weather.
ballistic
Refers to fruits that discharge their seeds forcefully; catapult fruits.
bamboo
Any of various usually woody and/or arborescent grasses of the tropical or temperate regions belonging to the genera Bambusa, Arundinaria, Phyllostachys, Sasa, Dendrocalamus, etc., most having hard, hollow stems.
banados
Shallow swamps in Paraguay.
Bancroft’s Law
A generalization that organisms and communities tend toward a state of dynamic equilibrium with the environment.
bank storage
Water absorbed by the bed and banks of a stream, returned in whole or part after ground water level falls.
bar

A continually moving deposit of sand forming a ridge along the seashore or the coasts of large lakes. 2. A unit of atmospheric pressure equivalent to 29.53 inches (750.1 mm) of mercury at 32 degrees at latitude 45 degrees.
barachore
A species in which the fruit or seed is desseminated by its own weight, e.g., walnuts, Juglans, falling to the ground. See also: autochore, diaspore.
barbed
Furnished with rigid points or short bristles, usually reflexed like the barb of a fish hook.
barbed trichome
A trichome with terminal or lateral retrorse projections, each projection being a barb.
barbellae
Short, stiff barbs.
barbellate
Finely-barbed.
barbulae
Outgrowths on the margin of the wings or in the throat of the corolla, sometimes with apical hairs or papillae (found in plants of the genus Scaevola.)
barbulate
Finely-bearded.
barchan
A distinctive isolated, crescent-shaped, sand dune with the ends projecting on the side opposite that from which the wind blows, common in Turkestan.
bare-root
Describes a plant that is prepared for transporting by removing all the soil around its roots.
bark
The outer covering of the trunks or branches of trees and shrubs.
bark-cambium
The layer of cells that produces new bark.
barogram
A continuous record of the readings of a barograph.
barograph
A self-registering barometer.
barotaxis
Response of an organism in response to barometric stimulus.
barotropism
Change of position of a plant or sedentary animal in response to a barometric stimulus.
barrens
A region where vegetation is absent or poorly developed.
barrier

Any feature or condition that restricts movement of organisms or prevents establishment of organisms which have migrated there. 2. A condition that prevents or significantly reduces crossbreeding of organisms.
barrier beach
A strip of land built up by the action of waves, currents and winds, and which protects inner areas.
basal (alt. basilar)

Growing from the base of a stem; used in reference to leaves at the base of the stem. 2. One of the main canes of a rose bush, originating from the bud union. 3. Describes cells at the base or insertion of the leaf, often of different shapes and colors from those of the main part of the leaf.
basal area (alt. basal cover, alt. ground cover, alt. cover)

The area of the cross section of a tree at a height of 4.5 feet above the ground, generally written as the total of the basal area of the trees in a forest in square feet per acre. 2. The surface of the soilactually covered by a plant, as compared to the full spread of the herbage, which in grassland ecologyoften measures at one inch above the ground.
base
The proximal portion of a structure; that part nearest the point of attachment.
base exchange capacity
A measure of the absorptive capacity of a soil for materials with exchangeable cations, a nonacid reaction. A soil with a high base exchange capacity will retain more plant nutrients and is less apt to leach than one with a low exchange capacity.
base flow
Springs; stream flow coming from subterranean sources in contrast to surface runoff.
base level
The lowest level to which a land surface can drop by action of water; the permanent base level is sea level.
base saturation
The proportion of the base exchange capacity that is saturated with metallic cations.
basidiomycete
A fungus-producing basidia.
basidium (pl. basidia)
The reproductive fungus cell (meiotangium) producing usually four spores on the outside.
basifixed
Attached by the base.
basin irrigation
An artificial method of watering plants in which a level field is surrounded by a ridge of earth so that a shallow body of water may accumulate before it soaks into the soil.
basin listing
A way of working land by constructing small dams at intervals across furrows to form basins for collecting rainwater, slowing runoff and erosion.
basionym
A specific or infraspecific name which has priority over other names later given to the same plant by different authors. See also: synonym, homonym, tautonym, autonym.
basipetal (adj. basipetalous)
Developing from the apex towards the base. See also: acropetal.
basiscopic
Facing the base of the axis on which it is borne.
basitonic
Describes flowering seasonal shoots which produce no leaves (except for bracts in some cases) below the inflorescence. See also: acrotonic.
basophilous
Refers to organisms which have adapted for life in alkaline soil or other medium.
bast
The fibrous portion of the inner bark.
bathyal zone (adj. bathyic)
The deep part of the ocean where light does not penetrate sufficiently for normal plant growth.
bathypelagic
Refers to deep parts of the ocean, but not including the ocean bottom.
bathyphyll
A modified leaf which attaches a plant to a substrate.
bathysphere
A pressure-resistant underwater structure with a spherical chamber in which scientists can descend deep into the ocean.
batten
Lath, often wired together as in snow fencing.
batten stripping
Plastic strapping which can be used in a manner similar to snow fencing.
bay (alt. embayment)
A notch in the shoreline, not restricted to any single form. It may be a lobe of water extending inland as deeply as a firth or fiord, or it may be as shallow as a quarter moon shape. See also: cove.
bayou
A creek or slow-moving stream.
beach
Where water meets land at the edge of lakes, oceans, etc. A barrier beach is a ridge of sandy deposits separated from the mainland by a section of water. See also: atoll.
beach pool
The barrier beach pool is a shallow lagoon formed inland from the barrier beach. 2. A sand spit beach pool is a shallow lagoon, generally S-shaped, that is inland from a sand spit, most often on the protected side of a headland.
beak
Prolonged narrow tip of the operculum.
beaked
Ending in a prolonged tip.
bearded
Bearing a long awn, or furnished with long or stiff hairs, as seen on the lower petals of some irises.
Beaufort scale
An artificial number scale invented by Francis Beaufort in 1805 to show approximate wind sp.edranging from 0 miles per hour (mph) for calm to 12 mph for a hurricane velocity in excess of 75 mph.
Beckmann thermometer
A thermometer for a narrow range at very accurate readings, such as one graduated to 0.01 degrees for a range of 7 degrees.
bed load
Rocks, sand, gravel, and other debris rolled along the bottom of a stream by high-velocity water, in contrast to silt load, which is suspended in place.
bed preparation
Mixing compost, ashes, and/or sand with natural soil to improve growing potential.
bedding plants
Plants used in large numbers in usually temporary displays. Most often annuals, tender perennials, and bulbs not able or intended to naturalize and which are grown indoors or under glass before planting.
bedrock
The layer of solid rock underlying a soil profile or other surface materials.
belly plants
A humorous term for inconspicuous plants, which often require botanists to crawl on their bellies to find them.
below
Refers to the position of one portion of an organ or plant in relation to another portion; the part “below” is the nearest to the attachment. See also: above.
belt
A long narrow area or strip of vegetation with characteristics which define it from adjoining areas.
belt transect
A type of vegetational analysis in which a portion of land a few inches or a few feet wide, and often a meter wide and 10 to 100 meters long, has constituent plants recorded.
bench mark
The point of reference in elevation surveys from where surveyors start to furnish an accurate survey.
bench terrace
A shelf-like embankment of earth, usually man-made, along a contour of sloping land to control runoffand erosion. See also: ridge terrace.
benthic
Refers to the bottom of any body of water, regardless of depth. Oceanic benthic division is composed of littoral, sublittoral, archibenthic, and abyssal-benthic zones.
benthos
Organisms which live on or at the bottom of oceans or fresh water, from the water’s edge down to the deepest water depths. See also: nekton.
bentonite
Absorptive and colloidal clay used as a sealing agent, as in lining ponds.
Berlese funnel
A device in which soil is placed, and heat and light are applied from above, forcing mites, collembolons, etc., into a container below it. See also: Baermann funnel, Tullgren funnel.
berm (alt. berme)

A narrow shelf, path, or ledge typically at the top or bottom of an escarpment or beside a road. 2. A mound or wall of earth.
berry (pl. berries)
A fleshy fruit that contains small seeds, the whole pericarp of which is fleshy or pulpy.
betacyanin
The nitrogen-based red pigments of beets, Beta spp., and other Chenopodiaceae, as well as flowers of Cactaceae, Portulacaceae, and others.
biannual
Occurring twice a year. See also: biennial.
biauriculate
Having two auricles.
biblical garden
A botanical area consisting of plants mentioned in the Bible.
bibracteolate
Having two bracteoles.
bicalcarate
Having two spurs.
bicallose
Having two callosities.
bicarinate
Having two keels.
bicarpellate
Having a two-celled fruit.
bicentric
Refers to a taxon which has two centers of dissemination or evolution.
bicolored (alt. bicolorous)
Having two colors on the same structure, often said of petals.
biconvex
Bulging outward on both sides, almost a sphere or orb.
bicornute
Having two horns.
bicostate
In mosses, having a double costa, which is usually much shorter than in leaves having a single costa.
bicrenate
Biserrate with both sets of teeth being shallow and rounded.
bidentate
Having two teeth.
biennial
A plant with a life cycle that is completed in two years or seasons, with the second season usually devoted to flowering and fruiting.
bifacial
Refers to leaves with distinct adaxial and abaxial surfaces.
bifarious
Growing in two ranks, such as needle-type leaves growing in two rows on opposite sides of a twig; distichous.
bifid

Cleft into two parts. 2. Divided into pinnae-bearing pinnules.
biflorus
Refers to a plant which flowers in both spring and autumn.
bifoliate
Having just two leaves.
bifoliolate
Having just two leaflets per leaf.
bifurcated
Forked or divided into two parts.
bigeminate
In two pairs; used in describing pinnate leaves which have only two pairs of pinnae.
bigeneric
A hybrid produced by crossing parents from two genera.
bilabiate
Two-lipped.
bilamellate
Made up of two plates.
bilateral symmetry
Describes flowers that can be divided into two equal halves by only one line through the middle.
bilocular
Two-celled.
binary fission
The division of a single-celled organism into two daughter cells.
binate
Growing in pairs.
binder
Hardened cement paste.
binder twine
Coarse jute or plastic cord for baling hay or tying bundles of wheat or other grain. See also: nursery jute.
binomial (syn. binary name, syn. binary combination)
A name consisting of two parts, the first being the genus and capitalized, the second being the speciesand usually lowercased initial with both names being italicized, e.g., Gypsophila elegans.
binomial nomenclature
The use of the genus and species names together to identify a given organism in the taxonomic system.
bioassay
Determination of relative strength of a substance by testing on an organism.
biocenose (alt. biocoenose, alt. biocoenosis)
All of the interacting organisms living together in specific habitat, usually containing producer, consumer, reducer, and transformer types. See also: ecosystem, community, association.
biochemical oxygen demand (abr. B.O.D.)
A test to detect and measure pollution in water by determining the quantity of oxygen already used up by oxidizable materials.
biochore
A subdivision of the biocycle which contains a group of biotopes which resemble one another; habitat. Examples are grassland, forest and desert.
bioclimate
Microclimate.
bioclimatic law (syn. Hopkins’ law)
The generalization that in temperate North America, weather-related events, such as bloom time, can be determined to be similar as you move up or down a mountain 400 feet, or one degree of latitude, or 5 degrees of longitude. In the spring, each of these measurements are eastward or upward and are four days later; in autumn, they are westward or downward and are four days earlier.
bioclimatology
The study of interrelationships between living things and the climate.
biocoenology (alt. biocoenotics)
The study of communities including qualitative and quantitative analyses.
biocycle
A subdivision of the biosphere, including saltwater, freshwater and land. Each consists of biochores. See also: biotope.
biodemography
Numerical and mathematical treatment of population problems.
biodiversity
The totality of genes, species, and ecosystems in a specified area, or the entire world.
biodynamic
Relating to a system of farming that uses only organic materials.
bioecology
A branch of biology treating interrelationships of both plants and animals among themselves and with their environment. See also: ecology.
biogenesis (adj. biogenic)
The principle that plants or animals can originate only from other plants or animals. See also: spontaneous generation.
biogeocenose
A specific ecosystem unique to a designated geographic area.
biogeochemical cycle
The circulation of chemical elements (e.g., oxygen, carbon, etc.) from the environment into plants and animals and back again into the environment.
biogeographic region
A biome.
biogeography
The branch of biology that treats the latitude and longitude of the location of plants and animals; range. See also: chorology.
biointization
The chemical treatment of seeds to stimulate growth.
biological clock
The rhythmic repetition of processes in organisms, like the need for sleep in mammals.
biological control
The use of living things to control pests, such as the control of aphids by lady beetles; integrated pest management.
biological efficiency
The ratio of the productivity of an organism or community of organisms to that of its supply of energy. A black bear would have a much higher efficiency hunting in the summer than in winter, so it is more efficient to hibernate during the winter months.
biological equilibrium (syn. biotic balance, syn. balance of nature)
The state of natural control, self-regulation of the numbers of plants and animals in a community, brought about by interactions within and between plants and animals and the effects of environmentsuch as weather. For example, as the numbers of white-footed mice, Paramyscus, rise, grass seeds, Graminae, decrease and the numbers of foxes, Vulpes, increase. See also: life cycle, pyramid of numbers.
biological factor
An influence that results from biological agents, including biotic factors such as lack of sunshine and physiologic factors like hormones. For example, death rates of white-tailed deer, Odocoilus, increase along highways during rutting season at night.
biological race (syn. biological strain)
A group of organisms which differ only in their physiological or ecological behavior from other groups in the same species, e.g., woodland white-footed mice, Paramyscus, run in straight lines where meadowwhite-footed mice run zigzag lines.
biological resources (syn. biotic resources)
The factors of biodiversity which are of direct, indirect, or potential use to humanity.
biological spectrum
A percentile tabulation of the plants of a community into the life form classes according to Raunkiaer’s classification.
biologics
Biological products such as vaccines.
biology
The study of living organisms.
bioluminescence (syn. phosphorescence)
The emission of light by living organisms such as fireflies and jellyfish.
biomass
The total mass of all living organisms in a given area.
biome (syn. biotic formation)
The ecology of a particular habitat, characterized by its unique plant and animal symbioticrelationships, and maintained by local climatic conditions.
biome-type (syn. biorealm)
A group of similar biomes, such as the temperate deciduous biome-type which includes all the deciduous forests of eastern North America, China and Manchuria, and Europe.
biometry
The statistical study of organisms.
bionomics
The study of the relationship of organisms to each other and their environment. See also: ecology.
bioregion
An area defined by social, biological, and geographic criteria, rather than geopolitical jurisdictions.
bioregional management (syn. bioregional planning)
Directional supervision over an area of similar habitat rather than by standard state/county lines, e.g., cooperative public, private, and business environmental planning for a major watershed such as Chesapeake bay.
bios
Plant and animal life.
bioseston
All living components of the seston.
biosphere
That part of the earth and its atmosphere that can support life.
biosynthetic (n. biosynthesis)
Describes a chemical compound produced by a living organism.
biosystem
Ecosystem.
biosystematics
The classification of living organisms that recognizes and differentiates biotic units into taxa on the basis of genetic relationships.
biota (syn. flora and fauna)
All of the living things, including animals, plants, fungi, and microorganisms, located in a given area.
biotechnology
Applied biological science, especially in genetic engineering and DNA technology.
biotic
Refers to any aspect of life, but especially to characteristics of entire populations or ecosystems.
biotic area (syn. biotic region)
A general term delineating any large area from adjacent areas on the basis of the composition of its biota.
biotic climax
The type of community capable of lasting for long periods of time under the present soil and climate conditions, in combination with the animals living there, including humans. See also: edaphic climax, physiographic climax, sere.
biotic environment
All living things found in the environment of an organism or community.
biotic factor (syn. biological factor, syn. biotic influence)
Environmental influences caused by plants or animals like shading or trampling. Sometimes used so nonliving effects are included, like landslides. See also: coaction.
biotic potential (syn. reproductive potential, syn. breeding potential)
The natural capacity of an organism to survive and reproduce, which is slowed or controlled by environmental resistance.
biotic pressure
The tendency of a species or community to extend its range.
biotic province
A major ecological section of a continent containing one or more regional communities of plants and animals. See also: biome, formation.
biotin

A substance which promotes or stimulates growth and repair. 2. One member of the Vitamin B complex.
biotope
A region environmentally uniform in conditions and in the flora and fauna which live there.
biotrophic
Describes an organism which cannot survive or reproduce unless it is on another organism.
biotype

A group of organisms sharing a genotype. 2. A particular physical characteristic distinguishing a population of an organism adapted to a particular environment which does not occur in populations of the same species in other environments.
biovulvate
Having two ovules.
bipartite
Divided into two parts almost to the base.
bipinnate
Twice pinnate; the primary leaflets being again divided into secondary leaflets. See also: bipinnatifid.
bipinnatifid
Twice pinnatifid. See also: bipinnate.
bipinnatisect (alt. 2-pinnatisect)
Refers to a pinnately compound leaf, in which each leaflet is again divided into pinnae.
bipolar distribution
Refers to a taxon present in both the northern and southern hemispheres with no apparent connection between populations.
Birge’s rule
A generalization theorizing that the thermocline is the transition layer in lakes in which the temperature decreases at a rate of one or more degrees for each meter in depth.
bisect
A line transect which shows the vertical and lateral distribution of roots at the walls of a trench with the above-ground parts of plants along the verge.
bisected
Completely divided into two parts.
biseptate
Having two partitions.
biseriate
Arranged in two rows.
biserrate
Refers to a serrate border where the principal teeth are serrated.
bisexual
A flower having both stamens and pistils. See also: synoicous.
bistratose
Having two layers of cells.
biternate
Twice ternate; with three pinnae each divided into three pinnules.
bitten
Praemorose, with ragged edges as though chewed.
bivalent
A pair of chromosomes, usually one from each parent.
bivalvular (alt. bivalvate)
With two valves.
bivoltine
Refers to organisms which produce two generations each year, like raspberries, Rubus, which produce fruit in late spring and fall.
black alkaline
Soil with a pH well over 7.5, covered with a dark crust of sodium or potassium carbonates.
black spot (alt. blackspot)
A fungus appearing as large, fuzzy black spots on rose leaves, especially in wet weather.
blacktop
Asphalt surfacing used for driveways, paths, and roads.
bladdery (alt. bladder-like)
Inflated, with thin walls like the bladder of an animal.
blade
The expanded portion of a leaf, frond, etc.
blanching
Excluding light from the green parts of plants so that they become white and tender; etiolated.
blast
Injury caused by disease or conditions such as hot winds that shrivel buds, branches, etc.
bleed
To have water seep to the surface of cement paste due to settling.
blend
Two or more colors which gradually merge but are distinguishable from a distance of six feet (used in describing dahlias.)
blight

A disease of plants resulting in withering, cessation of growth, and death of parts, especially young tissue. 2. An organism that causes blight, such as a bacterium, fungus, or virus.
blistered
Where the surface of an organ is puckered, the veins being tighter allowing the tissue to round up.
blizzard
A storm in which high winds drive fine snow and ice crystals, reducing visibility.
bloom

A flower or blossom. 2. A whitish, powdery, or waxy covering.
blossom
The flower, often applied to fruit trees.
blown

Past tense of to blossom; already opened, as in a full-blown rose. 2. No longer viable, perhaps from drought or disease, as “blown buds of barren flowers.”
blowout
A wind-blown excavation in loose soil.
blunt
Not pointed.
boat-shaped
Carinate, cymbiform, navicular.
bob-back
To severely dehorn or damage woody plants while attempting to prune, without understanding the process.
bog (adj. boggy)

An area of little or no drainage acidified by rotting vegetation, often with some open water surrounded by a floating mat of sedges, ericaceous shrubs, acidophilous species, and sphagnum mosses. A quagmire, which trembles or gives way underfoot. 2. Loosely, and not technically correct, a marsh, swamp, moor, fen, muskeg, or heath.
bog soil
A mucky or peaty surface horizon with a peat underlayment.
bole

The trunk of a tree below the first major branch. 2. The crisscrossed bases of palm leaves.
bolson
A depression lacking above-ground drainage in an arid or semiarid region, a term primarily used in Mexico and southwestern U.S.
bolt
To produce flowers and seeds prematurely.
bomb
A pressurized can of insecticide or other compound.
bonitation (alt. ecological bonitation)
The state of well being in a population, reflected by increasing numbers of individuals.
bonsai

The art of growing miniaturized trees or shrubs trained for aesthetic effect. 2. A plant grown in such a way.
bonsai scissors
A cutting tool with large handles and small, very sharp, blades.
bool and clay
A rough form of masonry using locally available, uncut stone and clay mortar.
border dike
Man-made ridges of earth to hold irrigation water within certain limits in a field.
border fork
A heavily-constructed garden tool that is shaped like a fork with round tines and is used for digging in garden borders and flower beds.
border irrigation
Flooding fields by the use of border dikes.
border strip
A demarcation surrounding a plot, usually given the same treatment as the plot.
bordered
Having a margin different from the rest of the leaf.
bore
A tidal wave with a forward wall of water three feet (one meter) or more in height, advancing upstream in a narrow river estuary.
boreal forest
A forest made up mostly of conifers, such as that reaching across North America from Newfoundland to Alaska.
boreal period
The climatic period from about 7500 to 5500 B.C., typified by warm dry conditions.
borer
An insect larva that tunnels into stems and trunks of shrubs, trees, etc.
bosket (alt. bosquet)
A small park or block of woodland with pathways, usually planted with small, understory trees, and often having statuary and waterworks.
bossed
With a conical protrubence rising from the center of a surface, like the bump on a scale of a pine cone.
bostryx (pl. bostryces)
A one-sided helicoid cyme.
botanical
Of or relating to plants or botany.
botanical garden (alt. botanic garden)
A garden devoted to the culture, study, and exhibition of plants.
botany

The scientific study of plant life. 2. A botanical treatise or study.
botrytis
A group of fungi that causes plant diseases. 2. The disease caused by such fungi.
bottom deposits
Those materials composing and overlaying the original basin or channel floor of a waterway.
bottom water
To pour water into a saucer underneath a container of a houseplant or jardiniere, allowing the soil to absorb the amount needed. This is often done on plants whose leaves are harmed by water spots, such as African violets, Saintpaulia spp. Generally any excess is poured off after an hour.
bottomland
Flood plain; rich deposits of loam left from flood water runoff.
botuliform
Sausage-shaped.
boulder clay
Unstratified intermix of clay and stones deposited by glaciers. See also: till, drift.
bouquet

Flowers picked and arranged in a bunch. 2. A distinctive fragrance.
brachyblast
A short branch or shoot.
brachypterous
Refers to organisms with short wings.
brackish

Somewhat salty, as water in saline soils or at the union of a river with the ocean. 2. Less correctly, nonsaline water that is distasteful or nauseous because it is stagnant, etc.
bract

A more or less modified leaf subtending a flower or belonging to an inflorescence, or sometimes cauline. 2. The similar structure in cryptogams surrounding reproductive organs.
bracteal
Of or pertaining to the bracts.
bracteate
Having bracts.
bracteolate
Having bractlets.
bracteose
With numerous or conspicuous bracts.
bractlet (syn. bracteole)
A secondary bract, as one upon the pedicel of a flower.
brambles
Plants of the genus Rubus, Rosaceae family, usually prickly and bearing fruit, e.g., raspberries and blackberries.
bran
The edible broken seed coats of cereal grain, often separated from the flour or meal by sifting.
branch
A secondary woody stem growing off of the trunk or main stem of a woody plant.
branchlet
Except for the twig, the youngest and smallest division of a branch.
breccia
Rock composed of angular, often crystalline, pieces in a matrix.
breeder
A person who applies genetics and other sciences in the orderly practice of improving a taxon.
bristle (adj. bristly)
A hair-like prickle.
broad base terrace
An embankment built to carry runoff water along a contour, usually with a rounded crown 15 to 30 feet (5 to 10 meters) wide, with gently sloping sides and a dished channel along the top.
broad-leafed (alt. broad-leaved)
Refers to a plant which does not have coniferous needles or grass-like leaves.
broadcast seeding
Sowing across large areas by scattering seed mechanically or by hand.
broadleaf (alt. broad-leaved)
Having broad leaves, rather than needle-like or scale-like ones.
broadleaf evergreen (alt. broad-leaved evergreen)
An evergreen plant that is not a conifer.
brochidodromous
Describes leaves with pinnate venation in which the secondary veins do not terminate at the marginsbut rather are joined in a series of prominent arches. See also: acrodromous, eucamptodromous, semicraspedodromous.
bromeliad
Any of the mostly epiphytic herbaceous plants of the family Bromeliaceae found primarily in the tropical and subtropical regions of the Americas, including the pineapple and Spanish moss.
broom-like
Having many parallel branches.
brown earth

Any of a group of intrazonal soils developed under deciduous forests. 2. Soils needing replanting or restoration, especially damaged wetlands.
brown forest soils
Those soils with dark brown surface horizons, but becoming lighter-colored beneath. They are rich in humus and are neutral to slightly acidic in composition, commonly appearing under deciduous forests that are quite rich in calcium or other bases. See also: brown soils.
brown podzolic soils
Those soils with thin layers of partly decayed leaves over a gray-brown layer of mineral matter and humus that covers yellow or yellow-brown acid B horizons. These appear under deciduous or mixed forests in cool, humid, temperate regions.
brown soils
Those soils with brown surface horizon, becoming lighter in color with depth. Calcium carbonate appears at one to three feet. These develop under grasslands and shrubs in temperate to cool semiarid climates. See also: brown forest soils.
Brownian movement (alt. Brownian motion)
A condition discovered by Scottish botanist, Dr. Robert Brown (1773-1858), in which rapid vibratory movement of microscopic particles holds the particles and others suspended in a fluid. See also: colloid.
browse

Food for livestock or wild members of the extended deer family, Cervidae, consisting of woody twigsand shoots, with or without leaves. 2. To feed on those plants.
browse line
A condition found in forests or brushland with an over population of browse animals like deer or goats, all branches and twigs eaten as high as the animals can reach.
brush

A thicket of shrubs, small trees, etc. 2. Branches lopped off trees.
brush matting

A covering of branches spread on eroded land to conserve water and lessen erosion that helps establishes trees and other permanent vegetation. 2. A covering of mesh wire along streambanks that holds brush in place to retard erosion.
brush pasture
An area with a natural cover of woody plants, set aside for foraging by goats or other animal browsers.
brushland
An area with shrubs and little other vegetation.
bryophyte
A plant in the botanical division Bryophyta, which includes mosses and liverworts.
bud
The rudimentary state of a stem or branch; an unexpanded flower.
bud eye
A dormant bud in the axil of a leaf, used to propagate through bud-grafting.
bud union (alt. budding union)
The junction on a stem, usually swollen, where a graft bud has joined the stock following the process of budding. Usually found at or near soil level.
bud-scale
A modified leaf, without lamina, protecting a bud.
budded
Grown from a bud grafted onto a desirable understock.
budding (n. bud graft)
Method of propagating woody plants. A cutting of one variety, called the scion, with bud(s) attached is joined onto another related species or variety, called the stock. As the tissues grow together a singleplant is formed with top-growth of the scion on the base of the stock.
buffalo wallow
An area in a grassland where large grazing animals have rolled, usually because of biting flies, killing the grass and losing soil.
buffer species
A plant or animal which may serve as an alternate food supply for a consumer animal, lessening the demand for a more desirable food species.
buffer strip
A transect of grassland or other erosion resistant vegetation located between or below cultivatedlengths or fields.
buffer zone

A strip that partially or fully encloses a study area or other designated area to protect the inner section from ecological disturbance by outside pressures; a transition zone between districts managed for different objectives, such as a wildlife preserve and a state hunting area.
buffering
Modification of environmental conditions by planting vegetation or changing topographic features. See also: reaction.
bulb
A short underground stem with fleshy scales or coats.
bulbate
Inflated.
bulbiferous
Bearing bulbs or bulblets.
bulbil (alt. bulbel, alt. bulblet)
A small bulb or bulb-shaped body, especially one borne upon the stem, and usually produced for asexual reproduction.
bulbous
Having the character of a bulb.
bulk density
The mass or weight of oven dried soil at 100 to 110 degrees Centigrade, per unit of bulk volume, including air space.
bullate
Blistered or puckered.
bunch grass
A style of grass that grows in tufts.
bundle cluster
A cluster such as that of pine needles.
bundle scars
Tiny, somewhat circular dots within the leaf scar, caused by the breaking of the fibrovascular bundles which run through the petioles into the blades of the leaves.
bundle sheath
A clasping collar surrounding a collection of similar parts such as the collar on pine needles.
bur (alt. burr)
A rough, prickly husk covering the fruit or seeds, such as the husk on a chestnut.
buried soil
One or more layers of soil, formerly at the surface, which have been covered by ash, sand, or other deposition.
burl
A deformed outgrowth on the trunk of a tree, often half a hemisphere in form, valued for the unusual wood grain; also the veneer made of such growths.
burn

Scottish word for brook. 2. The controlled process of removing dangerous buildups of combustible materials and destroying weed seeds, etc.
burn scar
A scar on a tree trunk left when tissues were damaged by fire. It may later be covered by new tissue.
bursicle
A pouch-like receptacle.
bush

A shrub, especially one that is low and thick with many stems rather than a single trunk. 2. A tree- or shrub-covered area in Australia or South Africa, or any uncleared land. 3. Sugar bush is a maple forestwhere sap is collected to make maple syrup or candy.
butte
A term from the western united States meaning an isolated hill with steep sides and a relatively flat top that is smaller than a mesa.
button center
A round center in a rose blossom, formed by unexpanded petaloids in very double roses.
buttress

A flange of tissue protruding from the trunk of a tree, tapering outward at the base. 2. A projectingstructure of masonry or wood for supporting or giving stability to a wall or building.
c horizon
In soils, the partly weathered rock fragments which are the parent materials for the upper A horizonand B horizon. This is occasionally lacking. See also: D horizon.
caatinga
The area of northeastern Brazil covered with thorn scrub.
cabling
Connecting tree limbs with steel cable to alleviate ice storm damage.
cactiform
With succulent stems similar to Cactaceae.
cactoid
Resembling a cactus.
cactus (pl. cacti)
A member of the family Cactaceae, having succulent stems and branches with scales or spines instead of leaves and found primarily in arid regions.
caducous
Falling off very early, usually applied to flower parts.
caingin clearing
A term used in southeast Asia for an area which has been cleared so that it can be used for farming.
cairn
Piled stones used as a landmark or memorial.
calcarate
Produced into or having a spur.
calcareous
Containing calcium or calcium carbonate (lime), as an alkaline soil.
calcareous ooze
Partially decomposed organic matter mixed with a quantity of calcareous material on the bottom of some bodies of water.
calcicole
A plant adapted to growing on limestone or alkaline soil.
calcification
A soil process where the surface soil is combined with calcium by the decomposition of plants, especially if a calcareous layer is formed. See also: podzolization.
calcifuge
A plant that does not grow well in lime or alkaline soil.
calcination
Decomposition due to the loss of bound water and carbon dioxide.
calcine
Like a calyx, or belonging to the calyx.
calciphile
A plant that must have lime or alkaline soil.
calciphobe
A plant preferring alkaline soils; an acidophilous plant.
caldera
A large crater formed by the collapse of the central part of a volcanic cone.
caliche

A crumbly crust of calcium carbonate formed on a stony base in arid climates. 2. A term used in Chile and Peru for deposits of sodium nitrate.
calicole
A plant living on chalky or limy soils.
caliper

An instrument used to measure the diameter of a tree, or other object. 2. The measurement thus attained, usually at breast height.
calli
Small outgrowths on the throat of the corolla of some plants, often acting as tactile guides for pollinators.
callosity
A hardened thickening.
callus (adj. callose)

A hard protuberance or callosity, sometimes resulting from a wound. 2. In the grasses, the tough often hairy swelling at the base or insertion of the lemma.
calmative
An herbal medicine that contains a mild sedative or has a calming effect.
Calvin cycle
The cycle of dark reactions of photosynthesis that occurs in the chloroplasts and involves the fixation of carbon dioxide and the formation of a six-carbon sugar.
calycine
Refers to a part of the calyx, or something that is like a calyx.
calycle (alt. calyculus)
The epicalyx, a second calyx or involucre beneath the true calyx.
calyculate
Having bracts around the calyx or involucre imitating an outer calyx.
calyptra
A lid or hood. In mosses, the thin veil or hood covering the mouth of the capsule.
calyptrate
Having a calyptra.
calyx
The outer part (perianth) of the flower, usually green and formed of several divisions called sepals, that protects the bud.
calyx bract
A bract which takes the form of a petal or sepal.
calyx lobe
The portion of the sepal which is apical, or remains free, in a fused calyx.
calyx tube (alt. calyx-tube)
A tube formed partly by the united bases of the sepals and partly by the receptacle. See also: hypanthium.
cambium
The internal layer of living cells between the inner bark and the sapwood where growth takes place that produces secondary xylem and phloem.
Cambrian

The oldest geological period in the Paleozoic era, about 500 million years ago. 2. Refers to organisms formed during that period.
campanile
A bell tower, usually freestanding.
campanulate
Bell-shaped; cup-shaped with a broad base.
campestral
Growing in fields.
campo cerrado
A Brazilian type of parkland composed of scattered trees in dense grass.
campos
Grassland located south of the equator in Brazil.
camptodromous
A pattern of venation where the secondary veins curve toward the margin without forming loops.
campylotropous
Describes an ovule or seed so curved as to bring the apex and base nearly together.
canaliculate (syn. tubulose)
Longitudinally channeled or grooved, especially on leaf stalks and midribs.
cancellate
A surface having the appearance of a lattice, with regularly arranged openings, e.g., the endostome of the Fontinalacae.
candle
The tender spring growth of the pine and some other needle-leaf evergreen species.
cane
A long, often supple, woody stem.
canescent
Hoary with gray pubescence.
canker
A patch of dead cells on a trunk or branches of a woody plant.
cano-tomentose
Density of hairs midway between canescent and tomentose.
canopy
The uppermost layer of branches and foliage of forest or a single tree.
canopy layer
The highest stratum of growth in a forest, where the trees form almost solid treetop canopies.
canopy trees
Forest trees which have reached a size where the crown becomes part of the uppermost layer.
cant hook
A term used in the united States for a tool used for rolling logs, consisting of a wooden lever with a moveable iron hook near the end, often with a lipped iron ring round the tip. See also: log lifter, peavey.
capability (alt. land capability)
A measurement of the suitability of land for some particular use without permanent damage to the land.
capacity formula
Mathematical rule used in hydraulics to calculate the extent or discharge volume of a channel.
capillary
Hair-like.
capillary action (syn. capillarity)
The phenomenon of a liquid spontaneously seeping up thin tubes due to adhesive and cohesive forcesand surface tension.
capillary porosity
The aggregate volume of small pores within a particular soil to retain water against the force of gravity.
capillary water
The part of soil water which is held cohesively as a continuous layer around particles and in spaces, most of it being available to plant roots.
capitate (syn. capitiform, alt. capitose)

Shaped like a head; collected into a head or dense cluster. 2. Terminated by a bulbous, swollen area.
capitellate

Shaped like a tiny head. 2. Clustered in a compact headlike grouping.
capitulum
A dense inflorescence of unstalked flowers shaped like a globe, such as a buttonbush, Cephalanthus; or sometimes shaped flat, as in Compositae.
caprification
The commercial process of pollination of fig plants by wasps.
capsid bugs
Green or brown bugs that suck sap of young growth, causing distortion of flowers and leaves.
capsular
Belonging to or of the nature of a capsule.
capsule

A dry dehiscent fruit composed of more than one carpel that splits partly open at maturity. 2. The enlarged distal end of the sporophyte; it contains the spores, and is sometimes known as the sporangium.
carapace
The hard case or shield covering part of the body of animals like turtles and some invertebrates.
carbohydrates
Neutral compounds of carbon, hydrogen and oxygen formed by green plants, including sugars, starches and cellulose.
carbon
Nonmetallic element found in all organic compounds.
carbon 14 dating
The selection and testing of undisturbed soils, buried materials such as wood, firepit remains, and other organic materials, for measurement of radioactive carbon 14 with a half-life of approximately 5,500 years, to determine the rough age of the selected materials.
carbon assimilation
Photosynthesis.
carbon cycle
The process of changing atmospheric carbon into sugar by photosynthesis in plants, synthesis of more complex organic compounds in higher plants and animals, and the return to carbon dioxide by respiration or death and decay of plant and animal tissues.
carbonate zone
A soil layer with concentrated carbonates, chiefly calcium carbonate, found most often in arid areas.
Carboniferous
Refers to the Pennsylvanian (upper carboniferous) and Mississippian (lower carboniferous) geological periods in the upper part of the Paleozoic era, about 200 to 260 million years ago.
carcinogen (adj. carcinogenic)
Substance capable of producing or inciting cancer.
cardinal points
The four chief directions of a compass reading: north, south, east, and west.
cardioactive
Herbal medicines that can affect the heart.
carina
Keel.
carinal
On or relating to a ridge or keel.
carinate
Having a keel or projecting a longitudinal medial line on the lower surface.
carminatives
Herbal medicines that can dispel gas from the intestines, relieving colic.
carnivorous plant
A plant subsisting on nutrients obtained from the breakdown of animal tissue, usually that of insects.
carnose
Fleshy.
Carnoy’s fluid
A liquid preservative for cytological materials, containing 3:1 absolute ethyl alcohol and glacial acetic acid.
carotene (alt. carotin)
Orange-yellow hydrocarbon, a pigment which commonly appears in such plants as carrots and squash; a precursor of Vitamin A.
carpel
A simple pistil or one member of a compound pistil.
carpid
Half of a schizocarp, e.g., one of the winged seeds of a maple.
carpogonium
The female organ of red seaweeds.
carpophore

The slender prolongation of the floral axis which in the Umbelliferae supports the pendulous ripe carpels. 2. The nonmotile spore of the carposporophyte.
carposporophyte
The post-sexual fruiting stage of a red seaweed.
carr
A fen.
carrying capacity
The maximum number of individuals of a particular species that a given area can maintain indefinitely.
cartilaginous
Hard and tough, but still flexible.
caruncle (syn. strophiole, adj. carunculate)
An excrescence or appendage at or about the hilum of a seed.
caryophyllaceous

Refers to members of Caryophyllaceae. 2. Refers to petals which have a long claw at the base.
caryopsis
A grain, such as grasses; a seed-like fruit with a thin pericarp adnate to the contained seed.
cassideous
Shaped like a helmet or hood.
castaneous
Of a chestnut-color; dark brown.
casual
A weed in a cultivated field which appears occasionally but does not naturalize.
casual species
Species which occur rarely in a community, such as a lilac, Syringa, but does not naturalize young plants in the area.
catabolism
Destructive metabolism releasing energy and breaking down complex materials in a living organism.
catadromous
When the first branch of a frond or vein in a pinnate leaf appears on the side facing towards the base.
cataphyll
A leaf, or leaf-like organ, usually found below ground.
catapult fruits
Those fruits that discharge their seeds forcefully; ballistic fruits.
catarobic
Refers to a wet or aquatic habitat where the slow breakdown of organic matter is occurring. Organic materials are given off into the medium with much of the oxygen used, but not enough to prohibit the occurrence of aerobic organisms.
caterpillars
The larvae of butterflies and moths, which often feed on leaves.
cathartics
Potent herbal laxatives.
cation
A positively charged ion.
catkin
A compact and often drooping cluster of reduced, stalkless, and usually unisexual flowers; an ament.
caudate
Having a slender tail-like appendage.
caudex

The persistent, sometimes woody base of an otherwise herbaceous stem. 2. A trunk, especially that of a tree fern.
caudicle
The thread-like or strap-shaped stalk of a pollinium.
caulescent
Having a manifest stem above ground.
cauliflory (adj. cauliflorous)
The state of flowering from the branches or trunk.
cauline
Belonging to the stem.
caustic (syn. corrosive)
Capable of destroying by chemical action.
cell
One of the minute vesicles of protoplasm consisting typically of nucleus, cytoplasm, and, in the plant cell, photosynthetic pigments, all enclosed by a cell wall. Any structure containing a cavity, as the cells of an anther or ovary, etc.
cell membrane (syn. plasma membrane, syn. plasmalemma)
A semipermeable limiting layer of cell protoplasm.
cell system pasturing
An attempt to replicate the method of natural grazing by bison. Livestock are placed in one section or cell of pasture with access to water until all vegetation is eaten or trampled to the ground, then moved to the next section. A common pattern features 10 cells with livestock moved monthly so each section is grazed in different months each year to encourage increase of ice cream grasses. See also: continuousgrazing, rotation grazing.
cellulose
The complex carbohydrate that is the principal component of cell walls of plants.
cement
A powder of lime and other minerals burned together in a kiln and finely pulverized which, when combined with water, hardens with hydration. It is used as the binding agent in mortar and concrete.
cenospecies
All the ecospecies that may exchange genes through hybridization, like the violets, Viola.
Cenozic
Refers to the geologic era extending from 40 million years ago to the present era, which started approximately a million years ago.
center of dispersal
The geographic area from which a taxon has spread or is spreading, such as where an alien plant was introduced.
center of diversity
A specific locality with high levels of genetic or species variance.
center of endemism
A unique geographic region with species which are known only to that area. See also: endemic.
center of origin
The geographic area in which a taxon originated and from which it is spread.
center pivot irrigation
A method of agricultural irrigation using a long, wheeled arm with many nozzles that pivots about the center of a circle; used primarily in arid regions.
central rays
The short immature ray florets comprising the central portion of the fully double flowers when at their prime stage (used in describing dahlias.)
central strand
The middle of many moss stems is made up of a bundle of much narrower and more slender cells, known as the “central strand”.; This is usually continuous with the midrib or costa of the leaves, similar to the vascular bundles in the higher plants.
centrifugal
Progressing or developing outward from the center or axis.
centripetal
Progressing or developing inward from the outside and towards the center or axis.
centromere
The point or region on a chromosome to which the spindle attaches during mitosis and meiosis.
centrosome
Transparent cytoplasm adjacent to the cell nucleus.
centrum
The central portion; used specifically for the large central air space in hollow stems, such as those of Equistetum.
cephalanthium
The flower head of a Compositae species.
cephalium
A woolly growth at the top of the stem of some cacti, such as Melocactus, on which the flowers appear.
cephalodia
Tiny thalli growing in the upper cortex of some lichens.
cereal
Grain or plants that produce grain–mostly from the Graminae family–used as food.
ceriferous
Wax-producing; waxy.
cernuous
Drooping or nodding capsule, somewhat inclined as opposed to erect.
certified seed
Seeds that have been approved by a legally recognized certifying agency as being qualified under established standards of germination; they are free from disease and weeds and are true to variety.
cespitose (syn. caespitose)
Growing in tufts; forming mats or tufts; often refers to a short plant with many stems or branches, forming a cushion appearance.
chafers
Beetles that attack plant roots as larvae and leaves as adults.
chaffy
Having or resembling chaff.
chain saw

A power tool for cutting timber. 2. A tool with interlocking individual teeth that are chained together, with ropes on either end and powered by muscle.
chalaza
The area of an ovule, opposite the micropyle, to which the funicle is attached.
chalk
Soft gray, buff, or white limestone formed from the shells of foraminifers.
channeled
Deeply grooved longitudinally, like a gutter.
chaparral
A dry land, xerophytic, formation of impenetrable thickets, composed of stiff, thorny, small-leaved shrubs.
character
An attribute or property of an organism, functional or structural, which may be adapted through environmental conditions within genetically determined limits.
characteristic diversity
The pattern of distribution and abundance of habitats and their species populations under conditions where human influence on the ecosystem is no greater than that of any other biotic factor.
chart quadrat
A cart or map of a sample area showing the placement and area of each plant.
chartaceous
Having the texture of writing paper or parchment and usually not green.
chasmogamy (adj. chasmogamous)
The opening of the perianth of a flower for the purpose of fertilization. See also: cleistogamy.
chasmophyte
A plant which grows in the crevices of rocks and rock faces. See also: chomophyte.
check dam
A short low dam constructed in a waterway to decrease the sp.ed of stream flow and to allow sediments to drop from the water.
chelate (n. chelator, n. chelation)
The process whereby an organic chemical bonds with and removes free metal ions from solutions.
chemical stratification
A condition found in temperate lakes during the summer and winter stagnation periods in which certain horizontal layers become different chemically from adjacent ones, often causing turnover. See also: thermal stratification.
chemoautotrophic
Being autotrophic and oxidizing an inorganic compound as a source of energy, as a chemoautotrophic bacteria.
chemolithotrophic
Describes an organism which obtains its energy from the oxidation of inorganic compounds.
chemoorganotrophic
Describes a heterotroph which oxidizes chemical bonds for energy but requires organic carboncompounds to grow.
chemosynthesis
A process found in certain bacteria in which nutrition is assured by the ability to oxidize inorganicmaterials.
chemotaxis
Movement of an organism caused by a chemical stimulus.
chemotrophic
Refers to an organism that produces energy from a chemical reaction which does not depend on light. See also: heterotrophic, autotrophic, phototrophic.
chemotropism
Movement or growth of cells or organisms in response to chemical stimuli.
chemotype
A population of plants belonging to a particular species that differs chemically from others of that species.
chernozem (syn. black earth)
A zonal group of soils with deep fertile surface soil, dark brown to black in color, rich in organic matter, grading into lighter colored soil below, and having a calcium carbonate layer at a depth of 1.5 to 4 feet (45 to 120 cm.). These are often found under tall grasslands in a temperate to cool, somewhat humidclimate.
chestnuts soils
A zonal group of soils with dark brown surface horizons grading into lighter colored soil below, and a calcium carbonate layer at depths of one to four feet (30 to 120 cm.). These are associated with grasslands in temperate to cool, subhumid to semiarid climates; moister than brown soils, drier than chernozem.
chianophile
A plant that can endure long snowy winters, or one that requires snow cover in winter.
chianophobe
A plant that does not tolerate long snowy winters, or one that can survive with little or no snow coverduring winter.
chiasma
A crosswise fusion, as occurs with chromosomes.
chicon
The blanched, compact shoots of chicory (Cichoricum intybus).
chilling requirements
Seeds which are dormant or doubly dormant may need actual outdoor climatic conditions of winter, or the indoor equivalent known as stratification. This is usually done by placing the seeds in moist soil in a plastic bag in the refrigerator for several weeks or months.
chimara (alt. chimera, alt. chimaera)
A plant formed of the tissues of two different species mingled together and being intermediate in characteristics between the two parents.
chiseling
Working deeply into the subsoil with a chisel plow to break compaction. See also: subsoiling.
chitin
Horny material that forms exoskeletons of insects, arachnids, etc.
chlorenchyma tissue
Parenchyma cell containing chloroplasts found in plants.
chlorinity
The chloride content of a solution, often compared to seawater which is 19.3.
chlorophyll
The green coloring-matter within the cells of plants.
chlorophyllose
Containing chlorophyll.
chloroplast
The microscopic body within the cell which contains chlorophyll.
chlorosis (adj. chlorotic)
A yellowing of the leaves, reflecting a deficiency of chlorophyll and caused by waterlogged soil or a lack of nutrients, often iron.
chlorotic
Refers to a plant that has chlorosis.
cholagogues
Herbal medicines which promote the flow of bile from the liver.
chomophyte
A plant that grows on rock fissures or crevices, on rocky ledges. See also: chasmophyte.
choripetalous
Refers to a corolla made up of separate petals. See also: polypetalous.
chorology
The study of regions or areas. See also: synchorology.
chresard
Water in soil that is available to plants for absorption.
chromatid
One of the usually paired and parallel strands of a duplicated chromosome joined by a singlecentromere.
chromatin
Material of the nucleus and chromosomes which can be stained deeply with certain dyes.
chromatophore

In plants, a plastid which contains pigment in a cell such as a chromoplast or chloroplast. 2. In animals, a cell or group of cells with pigment having the capability of color change.
chromogen (adj. chromogenic)
A pigment-producing microorganism.
chromoplast
A plastid other than a chloroplast, containing yellowish or red pigment.
chromosome
One of the set of bodies in the nucleus which determines the hereditary cell structure and function of a cell.
chrysalis
A pupa of a butterfly; broadly, any insect pupa.
Chrysophyta
A taxonomic division containing diatoms, golden-brown algae, and yellow-green algae, all single-celledand normally found in marine environments.
chubasco
A spinning disturbance near the Gulf of California, resembling dust whirls on land and waterspouts over water. These environmental disturbances reach great heights, becoming violent enough to capsize small craft.
chylocaulous
Refers to fleshy stems, like cactus.
chylophyllous
Refers to fleshy leaves, like agave.
ciliate
Marginally fringed with hairs (cilia).
cilium (pl. cilia)
A short, usually stiff, usually unicellular, marginal hair; a hair-like thread of the endostome, alternating with the segments.
cincinnal
Refers to scorpoid cymes and other curled floral arrangements.
cincinnus
A monochasial cyme on which flowers appear in an order along a spiral.
cinereous
Ash-colored.
cinnamomeous
Cinnamon-colored.
circinate (alt. circinnate)

Curved into a circle so that the apex is nearly or quite bent around to the leaf base. 2. Coiled in a spiral, with the apex at the center. 3. Coiled from the top downward, as the young frond of a fern.
circle of vegetation
All of the species and communities that are restricted to a natural vegetation unit.
circulus
One of the concentric circles on a fish scale.
circumboreal
Said of plants which surround the Northern Hemispere, appearing in both the Old and New Worlds, such as the dandelion.
circumpolar
Refers to species that occur all around the poles in either the north or south and in both the eastern and western hemispheres.
circumposition
A propagative method also known as air layering.
cirque
A deeply-eroded area with steep slopes in a region of past glaciers.
cirrate (syn. cirrhate)
Applied to leaves which curl up in drying. Cirrate leaves are more regularly curled than crispate leaves.
cirrhose
Having a wavy hair point.
cistern
An artificial reservoir for collecting and storing rainwater, as the type used for irrigation, which is often ornamental in design and frequently serves only as a decorative function.
citation
In botanical systematics, a quotation from a book or author referencing an authority or precedent.
CITES (alt. C.I.T.E.S.)
The Convention on International Trade of Endangered species, which provides regulations for the international trade in listed species of plants and animals.
citronella
A fragrant oil used in perfumery and as an insect repellent derived from the grass Cymbopogon nardus.
citrus
Members of the family Rutaceae, e.g., oranges and limes, etc., grown in warm areas as edible fruits.
cladocarporus
Having the sporophyte terminating a short special fertile branch; somewhat like half-way between acrocarpous and pleurocarpous.
cladophyll (alt. cladode)
A flattened, leaf-like photosynthetic stem or branch not itself leaf-bearing.
clambering
Describes a plant that sprawls or climbs but lacks tendrils.
clasping
A leaf whose base wholly or partly surrounds the stem.
classification
Systematic arrangement of hierarchal levels of taxonomy.
clathrate
Lattice-like; having thick lateral (adjacent) cell walls and thin surficial walls.
clavate
Club-shaped; gradually thickening from a slender base.
clay soil (syn. clay loam)
A soil, usually heavy and poorly drained, containing a preponderance of very fine particles.
clean cultivation
Hoeing out weeds and keeping the surface of the soil loose to lessen weed seed germination.
clearcut (alt. clear felling)
The cutting of all trees in a forest.
cleft
Divided nearly to the midvein.
cleistocarpous
Capsule opening irregularly, not by a lid or valves.
cleistogamy (adj. cleistogamous)
The condition of having flowers which self-pollinate in the bud, without the opening of the flower. See also: chasmogamy.
climax stage (alt. climax community)
The ultimate stage in the process of succession, occurring when a plant’s ecosystem has reached a point of stability.
climber
A plant that can climb given support.
clinandrium
In Orchidaceae, the part of the central column which holds the anthers.
cline (adj. clinal)
A gradual morphological or physiological change in a group of related organisms across their range, usually correlated to environmental or geographic transition.
clinker
The material that emerges from the cement kiln after burning. It is in the form of dark, porous nodules which are ground with a small amount of gypsum to produce cement.
cloche
Covers for individual plants, protecting against frost, birds and pests; a miniature cold frame.
clod
A lump of soil.
clone
A group of plants all originating by vegetative propagation from a single plant, and therefore genetically identical to it and to one another.
cloud forest
A mountain forest covered by a persistent mist that creates stunted trees and abundant epiphytes.
club root
A fungus that causes knobby roots in stocks, wallflowers, and other members of the cabbage family.
clump-forming
Describes a plant which reproduces vegetatively, forming smaller crowns around the parent which can be divided and planted in new locations.
cluster
A group of two or more occurring close together.
clypeate
Halberd or shield-shaped.
co-management
The sharing of authority, responsibility, and benefits among government, businesses, and local communities in the management of natural resources, e.g., cooperation among the National wildflowerResearch Center, Texas Department of Agriculture, landscape architects, and nurseries in the production of native plants for landscaping.
coalesce
To fuse together.
coalescence
The union of parts or organs of the same kind.
coarse-toothed
With large teeth; dentate, serrate.
cob
The rachis of the female corn spike, fruit of Zea mays.
cobblestone
A naturally rounded stone larger than a pebble and smaller than a boulder; especially such a stone used in paving a street or in construction.
coccus (pl. cocci)
A lobe of a schizocarp containing a single seed.
cochlear
Describes a coiled arrangement of the corolla lobes in a bud, a type of imbricate aestivation.
cochleariform
Rounded and concave like a spoon or ladle.
cochleate
Spiral, like a snail’s shell.
cocous (pl. Cocci)
One of the parts into which a lobed fruit with 1-seeded cells splits.
Codes of Nomenclature
The general rules for the assignment of scientific names to taxa.
codominant leaders
When two main branches of a tree are of equal strength and size, leading to the possible danger of the tree splitting.
coefficient of association
A mathematical statement of the frequency of occurrence together of two species not due merely to chance, calculated by dividing the number of samples in which both occur by the number of samples in which it would be expected they both would occur.
coensorus
In ferns, the extension of a sorus, or united sori that appear to be a single sorus.
coetaneous
Of the same age or existing at the same time.
coexist (n. coexistence)
To live together in the same place and at the same time.
coherent
Refers to parts that are usually separate but have become fused together, such as petals in a floral tube.
cohesion
The union of one organ with another of like nature.
coir
A stiff coarse fiber from the outer husk of a coconut.
cold frame
A frame covered in glass or plastic and without artificial heat used to protect plants and seedlingsoutdoors.
cold room
An area to keep plant specimens fresh until they can be pressed.
coleoptile (alt. coleophyll)
The first leaf following the cotyledon of a monocotyledon which forms a protective sheath around the plumule.
coleorhiza
Protective sheath around the radicle in a grass embryo.
collateral
Situated side by side.
collecting permits
Papers granting permission to collect specimens of flora of parks, nature preserves, foreign countries, or other protected areas.
collection bottles
Glass or plastic bottles with waterproof screw tops or vial type bottles, used to collect specimens such as pollen, buds, bark, seeds, insects, etc.
collenchyma
A type of supportive tissue consisting of elongated cells with thickened walls and containing chloroplasts.
colleter
A group or tuft of mucilaginous secretory hairs, often found near the base of the leaf lamina and on the calyx in memebers of the Apocynaceae and Asclepiadaceae families.
colliculate
Having small hill-like eminences.
colloidal clay
Extremely fine, microscopic particles of rock.
colloidal dispersion
A mixture containing particles larger than those found in a solution but small enough to remain suspended for a very long time. See also: colloidal suspension.
colloidal suspension
Minute particles remaining intermixed in a liquid without being dissolved. See also: Brownian movement, colloidal dispersion.
collum
The neck or tapering base of the capsule.
colonial
Forming colonies by means of underground rhizomes, stolons, etc.
colony
A group of lichen thalli growing together.
color charts
Hue chips used to simulate colors in the field, as flowers may lose or change color(s) when dried. Examples of such charts are the horticulture Society Color Chart, Nickerson Color Fan, and Horticultural Color Chart.
colporate
Describes a pollen grain which has both an elongated and a rounded aperture.
colpus (pl. colpi, adj. colpate)
An elongated aperture of a pollen grain with a length/breadth ratio greater than 2. See also: colporate, porate.
columella
In mosses, the central axis of the capsule; around it and between it and the outer wall of the capsule are borne the spores. Sometimes the lid adheres to it and is raised upon it.
column
In orchids, a structure formed by the union of stamens, style, and stigma.
column foot
In Orchidaceae, the basal platform to which the lip is attached to the column.
coma
Comal tuft, a tuft of leaves at the tip of a stem or branch.
combination
In taxonomy, the name below the rank of genus, which combines the genus name along with the speciesand lower ranks, such as subspecies, variety, etc.
commensalism
A type of symbiosis where there are two organisms from different species. One obtains food or other benefits from the other without damaging or benefiting it.
commercial synonym

An alternative choice of a legitimate name for a cultivar. 2. A shortened form of the original name, used where the original name is not conducive to sales, e.g., Ilex vomitoria. 3. A sales name chosen by the originator of the cultivar or with the originator’s approval.
commissure
The surface by which one carpel joins another, as in the Umbelliferae.
common name
Identifying term which often reflects appearance, legend, or use of a plant species. It may var. greatly by region, increasing the need for standard binomial nomenclature.
common property resource management
The supervision of a specific resource (such as a forest or pasture) by a well-defined group of users with the authority to regulate its use by members and outsiders.
comose
Furnished with or resembling a tuft of hairs.
compact
Pressed together or closely joined; in rhizomes, those with short internodes and closely spaced stipes.
companion plants

Plants which work well together aesthetically, perhaps because they have complementary coloring, or flower at different times of the year, etc. 2. Two or more plants that are used together because one or more are beneficial to one or more of the others, for instance by repelling pests that prey on the other(s).
comparative advantage
Relative supremacy with which a region or state may produce a good or service.
complete fertilizer
A compound containing the three essential minerals for growth: nitrogen (N), phosphorous (P), potassium (K).
complete flower
Having all the principal parts, particularly the stamens and pistils.
complex
A group of very similar and obviously related plants where it is difficult to find identification limitations, and where the relationships are not fully understood.
complicate
Folded upon itself; folded lengthwise.
composite
Member of the family Compositae, having compound flower heads, e.g., the daisy, the aster, the sunflower, etc. See also: compound flower head.
composite cultivar
A group composed of several closely related cultivars; grex.
compost
Decomposed organic maker, usually used to enrich the soil.
compound
Composed of two or more similar parts united into one whole. Compound leaf: one divided into separate leaflets.
compound flower head (alt. composite flower head)
A flower head with outer ray flowers forming petals surrounding the inner disc flowers, as in the Compositae.
compound leaf
A leaf divided into smaller leaflets.
compressed
Flattened, especially laterally.
concatenate
Linked, as on a chain.
concave
Depressed or hollowed out.
concavo-convex
Bulging out on one side and caving in on the other, like an eggshell or lens.
concentric
Having a common center, as the rings on a target.
conceptacle
A flask-like structure containing reproductive organs.
conceptual design
Rough drawing showing the general shape and location of all design elements such as buildings, fences, slopes, plantings, etc. in a landscape plan.
concolorous
Uniform in color.
concressant
Growing together, especially of parts that were originally separate.
concrete
A hard compact building material formed when a mixture of cement, water, sand, gravel, and perhaps other aggregates, undergoes hydration.
concretion
Mineral mass formed within another type of rock, often very different in appearance and composition.
conduplicate
Folded together lengthwise.
cone
A conical fruit consisting of seed-bearing, overlapping scales surrounding a central axis.
cone-scale
One of the scales of a cone.
conferted
Closely crowded. See also: congested, constipate.
confervoid
Formed of fine threads, as green algae.
conflorescence
A compound inflorescence consisting of two or more simple inflorescences.
confluent
Running into each other; blended into one.
conform
Similar in shape and size to others.
confused center
A flower center whose petals are disorganized, not forming a pattern.
congeneric
Belonging to the same genus.
congested
Close together. See also: conferted, constipate.
congregate
Growing in dense proximity.
conical
Cone-shaped; widest at the base and tapering to the apex.
conical root
A taproot.
conifer
A cone-bearing tree of the pine family, usually evergreen.
coniferous
Cone-bearing.
conjugation
The union of gametes.
connate
United; used especially of like structures joined from the start.
connate-perfoliate
Where opposite leaves are completely joined at the bases, leaving the appearance that the stem pierces through.
connective
The portion of a stamen which connects the two cells of the anther.
connivent
Coming into contact; converging, but not fused.
conoidal
Nearly conical.
conservation lists
Recommended protection lists of endangered, threatened, and protected plants, giving information from groups other than those with enforcement powers.
conservatory
A structure or room made primarily of glass and used for the cultivation of tender and exotic plants. A formal term for a greenhouse.
conspecific
Belonging to the same genus.
conspicuous
Easily visible without a lens, often extended to mean showy or prominent.
constricted
Used of capsules that become narrowed under the opening when dry.
construction documents
Final drawings with accompanying written specifications to be used by contractors in completing a landscape project.
consumers
Organisms which break down organic material (such as sugars and proteins) to obtain energy for their own growth, and then return inorganic components to the environment, where they are again available.
container gardening
Cultivation of flowers and vegetables in boxes, urns, garbage bags, etc., where direct soil planting is not feasible, as on apartment balconies, in penthouses and desert areas, etc.
container-grown
Grown from a seedling in the container it is to be sold in.
contiguous
In immediate contact.
continental
Describes a climate little influenced by sea, with large ranges between summer and winter temperatures.
continuous
Not interrupted.
contorted
Irregularly twisted or distorted.
contracted
Reduced in size by or as if by squeezing or forcing together.
contractile roots
Roots that can shorten themselves much like a worm does, drawing the plant down deeper into the soil. They usually have a wrinkled surface for expanding and contracting.
controlled burn
A fire started to rid an area of flammable materials in order to prevent a more dangerous or costly wildfire.
convariety (alt. convar)
A group of similar cultivars within a variable species or hybrids between two species; the term has now been replaced in most cases by the word “group”.
convergent
Coming in contact, but not fused.
convex
Arched outward.
convolute
Rolled up longitudinally.
coppice

A thicket or copse of small trees. 2. To cut a plant almost to the ground each year so as to produce more vigorous growth.
copse
A thicket of small trees or shrubs.
coralloid
Shaped like coral, with many branches.
cordate (alt. chordate)
Heart-shaped with the point away from the stem.
cordiform
Heart-shaped in all three dimensions.
cordon
A woody plant, most often a fruit tree, trained to grow as a single stem on a support.
coriaceous
Leathery in texture.
corm
The enlarged fleshy base of a stem, bulb-like but solid.
cormel (alt. cormlet)
A small corm that develops at the base of the mother corm.
cormophyte
A land plant belonging to the former botanical division Cormophyta, having a stem and root system.
cormous
Bearing corms.
corn fork
A fork shaped like a scoop shovel, but with tines instead of a blade.
corneous
Having a horny texture.
cornicle
Wax-secreting tubes of certain insects, such as aphids.
corniculate
Having horn-like projections.
cornucopia
Curved horn overflowing with edible produce, symbolizing abundance.
cornucopiate
Shaped like a cornucopia or horn.
cornute
Spurred or horned.
corolla
The petals of a flower; the inner perianth of distinct or connate petals.
corolline corona
Fleshy ridges or outgrowths of tissue attached to the corolla tube.
corona
A crown of appendages between the corolla and stamens, or on the corolla or stamens. See also: crown.
coroniform
Shaped like a crown.
corpusculum
The central part of a pollinarium, characteristic of the families Orchidaceae and Asclepiadaceae.
corrugate
Wrinkled or in folds.
cortex

In rhizomes and other axes, the tissue between the stele and the epidermis. 2. In lichens, the outermost layer of the thallus. 3. In seaweeds and fungi, the tissue external to the central tissue of longitudinal cells.
corticate
Having a cortex.
corticolous
Growing on the woody parts of trees and shrubs.
corymb
A flat-topped or convex open flower cluster, the outer flowers opening first. In the stricter use of the word, equivalent to a contracted raceme and progressing in its flowering from the margin inward.
corymbose
In corymbs, or corymb-like.
cosmopolitan
Widely distributed over the world.
costa
A rib; a midrib or mid-nerve of a pinna or moss leaf.
costapalmate
Ribbed or veined palmately.
costate
Having a costa. Ribbed; having one or more longitudinal ribs or nerves.
costule
An axis that is a branch of a costa or another costule.
cottage garden
A usually small, informal garden making optimal use of space.
cotyledon (syn. seed leaf)
One of the first leaves to appear after germination (there may be one, two, or more); the foliar portion of the embryo as found in the seed. See also: true leaf.
cotype
An additional type herbarium specimen from which the taxon is described.
counteradaptation
Reciprocal acclimation of two or more species to each other over a period of time.
counterirritant
Herbal preparations that produce irritation when applied locally to stimulate circulation in the area of another irritated area.
cove
A small bay, especially on inland lakes.
crateriform
In the shape of a saucer or shallow cup; hemispherical or more shallow.
creeper
A plant running along at or near the surface of the ground and rooting. See also: creeping.
cremocarp
A dry fruit consisting of two one-seeded carpels that separate into mericarps upon ripening; schizocarp.
crenate
Dentate with the teeth much rounded.
crenulate
Finely crenate.
crescentic
Shaped like a crescent moon.
crest
A fan-like appendage, as can be found on some fern fronds; in spores, a tall, irregular ridge.
cretaceous
Chalky.
cribose
Peristome teeth perforated with small apertures.
crinite
Bearded with long weak hairs.
crinoid
Like a lily.
crispate (syn. crisped)
Frizzled, curled, and twisted in various ways.
crisped
Curly-edged, as seen on some fern fronds.
crispy-hairy
With ringlets.
cristate
Bearing an elevated appendage resembling a crest.
crop rotation
Process whereby a different crop is planted in a field each successive year, preferably the following crop utilizing nutrients not greatly used by the preceding crop. Insect and disease vectors are also controlled by rotation.
crosier (alt. crozier)
The unopened coiled leaves of some ferns, especially on the dormant crown.
cross-fertilization
Fertilization by the joining of gametes from different individuals of the same species (cross-pollination), or from different species (hybridization).
cross-pollination
The transfer of pollen from the anther of the flower of one plant to the flowers of a different plant.
crotch
The angle formed between two joining branches.
crown

The site on a plant where roots join the stem. 2. In trees, the branches, twigs, and leaves that form the top of a tree. 3. The corona. 4. In roses, the region of the bud union; the point near soil level where the top variety and the understock are joined. 5. An inner appendage to a petal or to the throat of a corolla.
crown gall
Galls on shoots or roots of shrubs, especially those of the rose family, caused by soil bacteria.
crownshaft
An apparent extension of the bole in some Palmae, formed by overlapping and sheathing bases of the leaves.
cruciate
Cross-shaped.
crucifer (adj. cruciferous)
Any plant of the family Cruciferae, including the cabbage, radish, turnip, and mustard.
crumpled
Corrugate; wrinkled.
crushed granite
Pea-sized chips of granite used for paving.
crustaceous
Of hard and brittle texture.
crustose
A form of lichen lacking a lower cortex and rhizines and having thalli that grow in contact with the substratum.
cryogenics

The branch of physics dealing with the effects and production of very low temperatures, as applied to living organisms. 2. Dormant preservation by freezing, drying, or both.
cryptogam
A general name for plants, and plant-like organisms that lack flowers and are not reproduced by seeds, e.g., including ferns, mosses, fungi, and algae. The name is close in translation to something like “hidden marriage.”
cryptopore
A stoma which is immersed.
crystalline
Looking like crystals or having the nature of crystals.
cube (adj. cubical)
A solid with six equal square sides.
cubiform
Shaped like dice.
cucullate
Hooded or hood-shaped; cowled.
cucullate calyptra
In mosses, a calyptra that is hood-shaped and split on one side only.
cucullus
A hood-like tissue on some seeds.
cucumiform
Shaped like a cucumber.
cucurbit
Any of the various twining or climbing plants of the family Cucurbitaceae, including the melon, gourd, squash, etc.
culm
The peculiar aerial stem which bears flowers, found in grasses, sedges, and rushes.
cultigen
A plant developed in horticulture and found only under cultivation or as an escape.
cultispec
A cultigen that should be given species recognition.
cultivar
An unvarying variety of plant produced by selective hybridization, or, sometimes found in wild populations, and maintained by vegetative propagation or by inbred seed.
cultivar class
A taxon consisting of a group of cultivars.
cultivate
To work the soil in order to break it up and remove weeds.
cultivation

A term for the tilling of soil. 2. Working up the bare soil around plants to kill weeds and allow air and water to penetrate to plant roots.
cultivator
A tool which may range in size from a hand-pushed garden implement to sets of teeth attached to a tractor, for the purpose of digging weeds and rounding soil over the base of plants.
culton
A recognizable plant originating in or maintained by cultivation, reflecting that not all such entities must remain cultivars.
cultriform
Curved like a short, wide scimitar.
cuneate
Wedge-shaped; triangular with the acute angle downward.
cup
The cup-like involucre surrounding an acorn. See also: crateriform.
cupped form
Having an open center, with the stamens visible.
cupreous
With coppery appearance.
cupressoid
With foliage like cypress; Cupressus.
cupule
A cup-shaped anatomical structure, like that holding an acorn, which is made of hardened, joined bracts. See also: crateriform.
cupuliform
Cup-shaped.
cure
The hardening of concrete, often controlled by keeping it moist.
curvate
Flexing or bending from a straight line.
curvinerved
Having curved parallel veins, as the leaf of a dogwood.
cusp
A short, abrupt, rigid point. See also: cuspidate.
cuspidate
Tipped with a cusp or sharp and rigid point. See also: cusp.
cut flower garden
An area set aside for the growing of plants which are to be used for the production of flowers for arrangement inside the house.
cut flowers (alt. cutting flowers)
Species of flowers which are removed from the plant for arrangements and which easily retain their freshness.
cuticle
Skin or epidermis. The waterproof layer of the epidermis of plants.
cutin
A thin waxy covering on the outer layer of a leaf.
cutting
What growers of houseplants call slips, used for propagating new plants. The method involves cutting or breaking off a part of the plant, inserting it in growing medium so it can grow a new plant like the one from which it came. Hardwood cuttings are mature wood; softwood cuttings are taken in spring or early summer from tips when plants are actively growing; semihardwood cuttings are taken in late summer or fall when growth has slowed and wood is beginning to harden.
cutting garden
A garden of flowers which can be cut and used in arrangements.
cutwater
A V-shaped projection at the base of a pier or bridge support used to deflect the force of water in a river or stream.
cutworms
Soil-living brownish caterpillars that feed at night, often severing stems of herbaceous plants.
Cyanophyta
A taxonomic division containing cyanophytes, cyanobacteria, and blue-green algae. These may be singlecells or colonies, and reproduce by fission. They are found in many environments: damp soil and rocks, fresh water, and salt water.
cyathiform
Cup-shaped.
cyathium (pl. cyathia, adj. cyathiform, adj. cyathophorous)
Cup-shaped. An inflorescence with unisexual flowers surrounded by a cup-like cluster of involucralbracts, e.g., a poinsettia.
cycad
Any plant of the order Cycadales, consisting of palm-like, cone-bearing, evergreen tropical plants that reproduce by means of spermatozoids and have large pinnately compound, usually fan-shaped leaves.
cyclone (adj. cyclonic)

A hurricane or typhoon; winds blowing counterclockwise in the northern hemisphere and clockwise in the southern hemisphere, at sp.s of 90 to 130 miles per hour (150 to 217 km/hr) around a calm center, with a forward thrust of 20-30 mph (33-50 km/hr) and a width of 50 to 900 miles (83-1500 km) diameter. 2. Popularly, any storm moving in a circular motion like a tornado.
cygneous
Curved suddenly downward, like a swan’s neck.
cylindraceous (alt. cylindric, alt. cylindroid)
Rod-shaped.
cylindrical
Having the shape of a cylinder.
cymbiform
Boat-shaped.
cyme
A usually broad and flattish determinate inflorescence, i.e., with its central or terminal flowersblooming earliest.
cymose
Bearing cymes, or cyme-like.
cymule
A small cyme, usually sparsely flowered.
cystocarp
A carposporophyte contained in a wall of sterile filaments.
cystolith
A concretion within a cell cavity consisting of calcium carbonate and occurring in plants such as figs.
cytokinin
Any of various plant growth hormones, such as kinetin, that grow and promote cell division and delay the senescence of leaves.
cytology (adj. cytological)
The study of cells.
cytoplasm
The part of the protoplasm that is not the nucleus.
cytoskeleton
The network of protein filaments and microtubules in the cytoplasm that controls cell shape, maintains intracellular organization and, in some organisms, is involved in cell movement.
D horizon
The soil layer below the effect of weathering, made up of undifferentiated and unconsolidated parent materials, located immediately below the c horizon. See also: A horizon, B horizon.
dactyloid (alt. dactylose)
Finger-like.
dale
A vale or valley.
damping off (alt. dampen off)
Collapse of small seedlings due to fungi attacking stem at soil level.
dark reaction
Those chemical reactions in photosynthesis not requiring the presence of light and involving the reduction of carbon dioxide to form sugar. See also: Calvin cycle.
data retrieval
The process–manual or automated–of identifying and locating specific facts in stored documents, published or unpublished, and specimen collections of interest to particular uses. See also: literature retrieval, document retrieval.
day degrees
The mathematical statement of the sum of degrees above a threshold for a designated period, such as those above freezing for the growing season. See also: temperature summation.
day-neutral plant
A plant that blooms when the length of day is either long or short. See also: photoperiodism.
dbh
An abbreviation for the phrase “diameter at breast height”, a measurement of the thickness of a treetrunk about four feet from the ground.
deadhead
To remove the blooms after flowering to encourage the development of new flowers.
dealkalization
Removal of base ions from soil by leaching or by chemical treatment with exchangeable sodium. See also: alkali soil.
decaploid
Having ten sets of chromosomes.
decapod
Animals like lobsters and crabs from the order Decapoda, class Crustacea.
decidulignosa
Communities of deciduous trees and shrubs.
deciduous

Not persistent. 2. A tree losing its leaves at the end of the growing season; nonevergreen.
declinate
Bending downwards.
decomposer
Any of various organisms, usually a bacterium or fungus, that breaks down organic substances making the nutrients available again.
decomposition
The breakdown of tissues and compounds into simpler substances which act as soil nutrients.
decompound
More than once compound or divided.
decordicate
Lacking a cortex.
decreaser
A species that gradually disappears under continued grazing. See also: increaser, population density, cover.
decrescent
Becoming gradually less.
decumbent
Reclining, but with the tip ascending.
decurrent
Describes a leaf that extends down the stem below the insertion.
decurved
With a downward curve.
decussate
Alternating in pairs at right angles.
deep watering
Inundating an area with water for a long period of time, perhaps 24 hours or more, to permeate the deepest layers of subsoil, thereby pulling roots down where they will not easily perish from drought.
deferred grazing
The delay of grazing in forage after growth has started, until development has reached a stage that will promote additional vigor of the plants. See also: cell system pasturing, continuous grazing, rotation grazing.
deficiency disease
A stress condition or disease of an organism caused by lack of a nutritive substance such as a vitamin or mineral.
definite
Of a constant number, not exceeding 20.
deflected
Bent downward at a sharp angle.
deflexed
Bent or turned abruptly downward.
deflocculation
Soil aggregates containing clay separating into individual particles.
deflorate
Flowering has completed for the season.
defoliant
A chemical substance which causes a plant to drop its leaves.
defoliate (n. defoliation)

To cause the leaves of a plant to drop. 2. To remove the leaves of a plant.
deforestation
Clearing trees from a woodland, often done to promote food for wildlife as well as to provide timber.
deformed
Misshapend, distorted.
degradation
Loss of friability or fertility of soil resulting from leaching.
dehisce
To split apart and discharge seeds or spores.
dehiscence (adj. dehiscent)

Breaking open at maturity to discharge seeds or spores. 2. Opening regularly to let seeds or spores escape by valves, slits, etc., as a capsule or anther. See also: indehiscent.
dehorning
Severely pruning shrubs or trees, often carried to extremes of butchery.
delicate
Fragile; easily broken.
deliquesce (n. deliquescence, adj. deliquescent)

To branch into many subdivisions and lacking a main axis. 2. To liquefy or become soft with age, as occurs with some fungi.
deltate
Broadly triangular with an obtuse apex.
deltoid (alt. deltate)
A low triangle attached at the middle of the wide part, rather than at the point; shaped like the Greek letter delta.
demarcation
An outline, boundary, delimitation, separation.
deme
One or more populations of a taxon; a population that will interbreed.
demersed
Constantly under water.
demography
The study of populations and their analysis regarding reproduction, deaths, age, etc.
demulcents
Herbal medicines that can break up phlegm or other mucus; sometimes used externally to sooth and soften skin.
dendroid (alt. dendritic)

Shaped like a tree. 2. Arranged in a shape like a tree, like some hairs on Cruciferae.
dendrology
The study of woody plants.
denitrification
The loss or removal of nitrogen or nitrogen compounds; specifically, a reduction of nitrates or nitrites caused by bacteria that usually results in the escape of nitrogen into the air.
density-dependent factor
An influence that hinges on a certain density of individuals to be fully effective, like peach trees or Amygdalis that must be 75 feet or closer for insects to successfully cross pollinate them.
density-independent factor
An influence that is effective regardless of density of a population, like the influence of a drought. Some scientists dispute the existence of such a factor. See also: nonreactive factor.
dentate
Toothed, usually with the teeth directed outward.
dentation
The teeth along the margin of an organ.
denticle
A small tooth; in plants of the family Rubiaceae, thick papillate tubercles along the margin of the interpetiolar stipules.
denticulate
Minutely dentate.
dentiform
Tooth-shaped.
deoperculate
Applies to a capsule after its lid has detached.
deoxyribonucleic acid (abr. DNA)
The chief material constituting genes and chromosomes.
depauperate
Starved; prevented from coming to its natural size through lack of nourishment.
dependence
An association between organisms in which the dependent one receives benefits but gives no benefit to the other, characteristic of such plants as mistletoe, Phoradendron, living on an oak, Quercus; not reciprocal. See also: symbiosis, competition, coaction, parasite.
dependency zone (alt. dependency range)
A belt surrounding an area of private land which supplements the use of the private land with additional use of public land.
dependent property (alt. dependent range)
Privately controlled land or water assumed to have special claim for use of public or cooperatively controlled range.
deplanate
Flat.
depressed
Somewhat flattened from above.
desalinization

The process of removal of salt from sea water to make drinking water. 2. Removal of salts from soil, usually by leaching.
descending
Bending gradually downward.
description
A list of characters which gives the attributes or features of a specific taxon.
desert grassland
Areas of the southwestern united States characterized by several species of adapted deep-rootedgrasses such as grama grass, three awn grass, and curly mesquite.
desert pavement
The stony surface of dryland areas when fine materials have disappeared from actions of water and wind.
deserta
Various types of plants found in areas that are low in available water like arid deserts, saline deserts, permafrost deserts, tundra, strand vegetation, and rocky mountainsides.
desiccate (n. dessication)
To dry up.
desilting area (alt. desilting basin, settling basin)
An expanse set aside above a dam, pond or field solely for the deposit of silt and debris from flowing water. See also: detention basin.
detention basin (alt. detention dam, detention area, detention pond)
A man-made holding area for the purpose of storing sewer overflow and surface runoff until such time as the water may be released for treatment without causing flooding.
determinate
Having growth where a bud or flower terminates the growing tip, e.g., a determinate inflorescence.
dethatcher (alt. de-thatcher)
A tool which combs clippings and thatch to the surface of the lawn.
detritivores
Organisms such as fungi, bacteria, and earthworms which consume nonliving material.
detritus

Any loose material that has separated from a mass, such as gravel from a rock face.
Devonian
A geological period in the Paleozoic era, which lasted from about 325 million years to 45 million years ago.
dew point
The temperature at which air is capable of holding no additional water vapor, with either a drop in temperature or an increase in water vapor causing the vapor to condense into liquid as rain, fog, or dew. During the course of precipitation, the relative humidity is 100 percent and the saturation deficit is zero.
dextrorse
Turned to the right.
diagonal
The average, a compromise of position.
dialypetalous
With separate petals.
dialysis
Separation of dissolved substances by their unequal diffusion through semipermeable membranes.
diameter at breast height (alt,. diameter breast high, abr. d.b.h., abr. DBH)
The width of the trunk of a standing tree, measured at 4.5 feet (1.3 meters) above ground surface.
diandrous
With two perfect stamens.
diaphanous
Very thin and transparent or translucent.
diaphoretics
Herbal medicines that promote perspiration and lower temperature.
diaphragm
Dividing membrane or partition.
diaspore
A reproductive portion of a plant like a seed or bud, that is dispersed and may give rise to a new plant. See also: disseminule.
diastrophism
Displacement of the earth�s crust by folding or slippage, causing the formation of mountains, chasms, etc.
diatom
Any of the minute planktonic unicellular or colonial algae of the class Bacillariophyceae, which contain silica.
diatomaceous ooze
Material consisting of siliceous remains of diatoms, found in cold seas.
dibble (alt. dibber)
A small hand implement used to make holes in the ground for plants, seeds and, most often, bulbs.
dibrachiate
Refers to branches that spread widely.
dicarpellary
Composed of two carpels.
dichasium
A cyme with opposite branching below the terminating flower.
dichlamydeous
Describes a flower which has two whorls of perianth parts.
dichloro-diphenyl-trichlorethane (abr. DDT)
An insecticide extensively used during the period after World War II that is now outlawed in the unitedStates and most of the industrialized world.
dichogamy
The differing times of maturation of stamens and pistils in a flower.
dichotomous
Forking regularly by pairs.
dichotomous key (syn. floral key)
A series of choices leading to the identification of a species.
dichotomy
The division of a growing point into two halves.
diclinous (alt. diclinic)
Having the stamens and the pistils in separate flowers.
dicotyledon (adj. dicotyledonous)
A flowering plant with two cotyledons.
dicoumarol
A compound derived from spoiled sweet clover hay and used to delay clotting of blood.
didymous
Twin; found in pairs.
didynamous
Refers to stamens found in two pairs of unequal length.
dieback
The dying of the outer portions of a plant due to disease or weather damage; death of part or all of the woody portion of a plant.
diel
The 24-hour period of night and day. See also: diurnal, nocturnal, crepuscular.
differential species (alt. differentiating species)
A species with high fidelity to a particular community that can be used to distinguish vegetation units.
differentiation

Maturation of a cell, organ, or immature organism from juvenile to adult status. 2. The development of new kinds of organisms in the course of evolution.
difform
Not similar.
dig in
To bury during the process of turning soil for planting, as occurs with compost, fertilizer, etc.
digamous
With both sexes in the same flower cluster.
digestion
The conversion of complex organic substances into simpler fats, proteins, and carbohydrates by chemical reactions and enzymatic processes.
digitalin
One of the mixtures of glycosides used in making digitalis.
digitalis
A drug derived from the seed and dried powdered leaves of plants of the genus Digitalis containing important glycosides and serving as a powerful cardiac stimulant and a diuretic.
digitate
Finger-like; compound, with the members arising together at the apex of the support.
digonous
Two-angled, as are the stems of some cacti.
digynous
With two separated carpels or styles.
dihybrid
A crossbreed; an organism that came about from the breeding of parents that differ in at least two characters, e.g., leaf shape and plant height.
dike
An embankment built to protect land from flooding or erosion, or to stop the loss of water from wetlands.
dilettante
A rather derogatory term for one who becomes interested in one field, then another, never delving deeply into any of them, an amateur, or at best a generalist.
dimerous
Having all the parts of the flower in twos.
dimidiate
Halved diagonally, or as if one half was wanting.
dimorphic
Having two shapes; in ferns usually refers to fertile versus sterile laminae or portions of laminae.
dimorphism
The state of a plant or animal showing two forms or colors in the same population, e.g., snowshoe rabbits with winter white and summer brown coats. See also: polymorphism.
dimorphous
Occurring in two forms.
dioecious (alt. diecious)
Unisexual, with male and female flowers on separate plants.
dip
A depression, as between two sand ridges.
diplecolobous
Describes cotyledons in a seed which are twice folded transversely.
diplochory
Describes seed dispersal involving two or more modes. For instance, a bird eats a fruit and disperses seed in it’s droppings and these are later carried away by insects.
diploid
Having two sets of chromosomes.
diplostemonous
The stamens of the flower are arranged in two separate whorls, the outer one with stamens alternatewith the petals, the inner one opposite the petals.
direct seeding
Planting straight into soil where the plant is to grow rather than germinating seeds indoors and transplanting.
direct sunlight
Unobstructed sunlight. See also: skylight.
dirt
Soil.
disarticulating
Breaking apart at the joints when mature.
disbudding
The removal of the side flower buds around a central bud to induce extra-large blooms; or to selectively remove buds to conserve strength in a newly-transplanted perennial.
disc floret
One of the small tubular, actinomorphic florets which make up the central part of the flower head in Compositae, each with a pistil and stamens but generally no other conspicuous flower parts. See also: ray floret.
disciform
Round and flattened. See also: discoid.
disclimax
A one-time climax community which has been disturbed by people or their domestic livestock, such as a deciduous forest being replaced by a cropland.
discoid (adj. discoidal)

Resembling a disk. 2. In Compositae, a flower head without ray flowers, having disk flowers only.
discolorous
Refers to a leaf which has different colors on the two surfaces.
discontinuity (alt. disjunction)
A gap in the range of a taxon. See also: distribution, disjunct.
discrete
Separate, distinct.
dish garden
A group of plants growing in a bowl or a shallow pot.
disjunct
Separated geographically, especially the population of a restricted area lying outside a main range.
disk
A development of the receptacle at or around the base of the pistil.
disk flower (alt. disc flower)
In Compositae, the tubular flowers of the head, as distinct from the ray.
disk harrows (alt. disk)
A field implement with disk-shaped blades which break up clods and level soil.
disoperation
Interaction between organisms in which one or all are harmed, as competition of maple seedlings, Acer, results in tall weak growth.
disparate
Unequal, not similar.
dispersal

The actual passage of disseminules or organisms from one place to another. 2. The history of the movement of a group of organisms, such as the introduction of zebra mussels into the Great Lakes and down the Mississippi River. See also: migration, establishment, spread.
dispersion

The pattern of distribution of individuals within a population, especially in prediction of probability. 2. In soil, the breakdown of aggregates, resulting in a single grain structure. Generally the more easily the soil disperses, the more it will erode.
disphotic zone
The depth in bodies of water where light is inadequate for photosynthesis but satisfactory for animal life.
dissected
Cut or divided into numerous lobes or divisions; a deeply cut leaf, the cleft not reaching to the midrib.
dissemination
The process by which propagules or organisms are spread.
disseminule
A plant part that can be easily separated from the parent plant, is dispersed, and can grow into a new plant. See also: diaspore.
dissepiment
A partition in an ovary, pericarp, or fruit.
distal
Towards the apex in position.
distant
Similar to parts that are separated and not overlapping.
distichous
In two vertical ranks.
distinct
Separate; not united; evident.
distributary channels
A system of forking streams flowing away from the main stream and not returning to it, as occurs in a delta or on an alluvial plain.
distribution

The geographic range, continuous or discontinuous, of a plant, animal, or community. 2. Dispersalpatterns of occurrence of individuals in a taxon from a certain area, such as a) random distribution, poisson distribution, normal distribution; b) nonrandom above normal distribution, contagious dispersal, over-dispersion, hyperdispersion; and c) nonrandom below normal distribution, hypodispersion, or even-spaced distribution. The terms following each letter are synonyms.
distylous
Refers to a flower with two styles.
dithecal
Having two pollen sacs or cells.
diuretics
Herbal medicines taht can promote the flow of urine.
diurnal
Describes flowers which only open in daylight.
divaricate
Widely divergent.
divergence
The circumstance in which seres of similar origin become less like each other as succession moves toward climax. See also: convergence.
divergent
Buds which point away from the twig.
diversion dam
A barrier constructed for diverting part or all of the water in a streambed into a different watercourse.
diversity index
The mathematical statement calculated by dividing the number of species in a specified area by the number of individuals of all of these species.
division

A major taxonomic grouping, ranking just below kingdom and above class. In the animal kingdom, it is usually replaced by the phylum. 2. Method of propagation for clump-forming plants through pulling apart fibrous-rooted clumps. 3. The removal of suckers from a parent plant for the purposes of propagation.
divisural line
The line down the teeth of a peristome, through which they split.
document
In botany systematics, this term can refer to herbarium specimens and equipment, as well as to the standard reference to written materials.
document retrieval
A more comprehensive term than ‘literature retrieval’ since it encompasses unpublished materials such as manuscripts, research notes, films, specimens, as well as published literature. See also: data retrieval.
documentation system
In botany systematics, this term covers the total strategy for identifying, citing, referencing, substantiating and authenticating, validating, verifying, and vouchering information used for a specific purpose in a taxonomic study.
dodder
Any of the annual parasitic wiry twining vines of the genus Cuscuta that lack chlorophyll and have tiny scales instead of leaves.
doldrums
The equatorial belt lying between the two trade wind belts that is characterized by calm or light variable winds and low atmospheric pressure.
doleiform
Shaped like a barrel.
dolomite (adj. dolomitic)

A mineral consisting of a calcium magnesium carbonate. 2. Limestone or marble rich in magnesium carbonate.
domatium (pl. domatia)
A small structure located in the axils of the primary veins on the lower surface of leaves in some woodydicotyledons, usually consisting of depressions and being partly enclosed by leaf tissue or hairs.
dominant species
A species that exhibits ecologic dominance or social dominance over adjacent ones. See also: secondary species.
donga
A South African term for a gully with steep sides or a dry watercourse.
dormancy
Temporary cessation of growth.
dormant
In the state of suspended growth.
dormant oil (syn. dormant spray)
Viscous solution containing pesticides and/or fungicides and applied to trees and shrubs in late fall and winter while the plant is dormant.
dorsifixed
Attached at the back.
dorsiventral (alt. dorsoventral)
Having distinct back and front (or upper and lower) surfaces, or placed with reference to the back or front.
double (alt. double-flowered)
Having more than the usual number of petals, often arranged in extra rows.
double citation
The occurence of two names at the end of a taxon, the first name, in parentheses, identifies the authorwho named the plant originally. The second name CITES the author of the new name.
double digging (syn. double trenching)
A method of inverting soil where the richer, top layer of soil of one trench is placed in the bottom of a subsequent trench and covered with the soil of a lower level.
double fertilization
An exclusive process of angiosperms in which one male nucleus pollinates the egg nucleus to form a zygote, which develops into an embryo, while the other male nucleus joins with two other nuclei in the embryo sac to form endosperm, which can be found in corn, Zea mays, and other grasses.
double-toothed
Each tooth bearing smaller teeth.
double-worked
Twice grafted. The plant consists of the rootstock, an intermediate scion, and the upper scion.
down
An undulating, usually treeless upland plain having sparse soil.
downland
An Australian term for temperate grasslands.
downy
Covered with fine hairs.
downy mildew
A fungus forming grayish downy patches, usually on the underside of leaves.
drainage basin
Any of the largest natural subdivisions of the watersheds of a continent, such as the Mississippi, Columbia, and Colorado basins of North America.
drainage terrace
A graded embankment constructed with a relatively deep channel and low ridge, primarily for the runoff discharge from a hillside.
drawdown
The process of partially or completely lowering the water from a wetland with pumps or other mechanical devices. The purpose of drawdown is to manage vegetation and wildlife.
drepanium
A floral cyme shaped like a sickle.
dried flowers
Desiccated blossoms which retain shape and color over a long period of time, especially for winter arrangements.
driers
Blotters, newsprint or other materials which absorb moisture from plants in plant presses.
drift barrier
An open structure, like a wire fence, constructed across a stream channel to catch driftwood.
drift fence
A fence constructed to prevent livestock from wandering from their allotted range to another one, often used as an extension of natural barriers such as large rivers and cliffs.
drift ice
Parts of icebergs in the open sea beyond the areas of pack ice.
driftwood
Any wood from trees or structures which has floated in water.
drill seeding
Sowing seeds with an agricultural implement which makes furrows into which it drops seeds. See also: broadcast seeding.
drip irrigation (syn. bubbler irrigation)
A method of irrigation where tubes or hoses, sometimes porous, are brought close to plants and water is allowed to trickle out at a very slow, but constant rate.
drip line
The line that could be drawn on the ground under a tree beneath the outermost tips of the branches. Rain flows off the tree at this point, so it is the area where roots congregate and the best point to place fertilizer, water, etc.
drip point (alt. drip-point, alt. drip-tip)

A leaf tip with an extension–acuminate, caudate, aristate–from which water drips during wet conditions. 2. A long drooping tip on leaves, particularly those of rain forest trees.
drip zone
The area around the base of a tree that lies below and within the circumference of the branches, etc., which forms its crown.
drooping
Erect at the base, but with the top part bending downward.
drought
A period of dryness; especially one that causes extensive damage to plants.
drought resistance (n. drought resistant)
The capability of an organism to survive extended dry periods with little or no injury. See also: xeric.
drought tolerant
Capable of surviving for extended periods with little or no rainfall.
drumlin
An elongated or oval hill made up of glacial drifts, usually compact and not stratified, and often with the longer sides parallel to the movement of the glacier when the soil is dropped.
drupaceous
Resembling or of the nature of a drupe.
drupe
A fleshy or pulpy fruit with the inner portion of the pericarp (1-celled and 1-seeded, or sometimes several-celled) hard or stony.
drupelet
A diminutive drupe.
dry farming

Agricultural operations in semiarid or arid regions without the use of irrigation. 2. A system of cultivation making heavy use of mulch and fallow periods to absorb and retain much of the rainfall.
dry stone
Building in stone without mortar.
dry wall
A wall built without the use of concrete, e.g., a stone wall.
drying frame
A structure, usually collapsible for easy transport, which holds plant specimens open to air and natural or artificial heat for rapid drying, thus preserving color and preventing mildew.
duckfoot
An agricultural implement with horizontal V-shaped blades that cultivate just below the surface of the soil without turning it over or burying crop residues.
duff
The partially decayed leaves, branches, etc., on a forest floor.
dune
A generally unstable mound or ridge of sand built up by winds, most common in deserts and along shores.
dune grass
Various species of Graminae with long roots, adapted to living on dry sand and used to stabilize dunes.
dune sand

An area of tiny rock particles accumulated by wind action into mounds or hummocks, generally with little or no vegetation and undergoing continual redeposition. 2. Refers to rock particles with diameters of 0.1 to 0.4 mm. which have been heaped up by the wind, even if found far underground.
dung
Feces, animal droppings, scat.
duplicate
Folded twice.
duriherbosa
Vegetation that has perennial roots, but whose tops die back each winter, like those of grasslands.
durilignosa
Woody perennial vegetation with broad hard leaves, like chaparral.
dust

n. A powdered chemical poison to kill insects or disease. 2. v. To apply the powder.
dust devil (alt. dust whirl, syn. remolino)
An intensely whirling column of air caused by a bubble of hot air caught between two cooler layers of air. Dust devils range from a foot to a mile high (30 cm. to 1.5 km.) mostly in arid regions; they spin dust and debris but can also shake houses and slam birds into trees.
dust mulch
Keeping the surface of the soil from caking by repeatedly working it loose, preventing the germinationof weed seeds.
Dutch elm disease
A fungus spread by bark beetles that causes wilting and dieback on elms.
dwarf

A plant that, due to an inherited characteristic, is shorter or slower growing than normal forms. 2. Dahlia varieties which normally do not produce plant growth over 24 inches in height.
dynamic equilibrium
A system that retains much the same condition because of the action of opposing forces which proceed at more or less equal rates. See also: balance of nature.
dysgenic (alt. eugenics)
Refers to detrimental influences on the genetic properties of a community or population.
dystrophic

Relates to or is caused by faulty nutrition. 2. Refers to a lake with high humus material, sparse bottom fauna, and low dissolved oxygen.
earth anchors
Steel pegs with semicircular bases to act as supports for larger trees, tying animals, etc.
earthworm
Any of the terrestrial annelid worms of the class Oligochaeta, especially those of the familyLumbricidae, that aerate and enrich soil.
earwigs
An insect that makes ragged holes flowers and leaves.
ebeneous
Black as ebony.
ecad (syn. ecophene, syn. environmental form)

A habitat form. 2. An organism showing somatic adaptations to a certain environment that are not hereditary. See also: phenotype, ecotype.
ecallose
With no callus.
eccentric
Not located at the geometrical center.
ecesis
The establishment of a plant or animal in a new habitat.
echard
Soil water not available for absorption by plants.
echinate
Beset with prickles.
echolocation

Acoustic orientation such as that used in sonar. The ability of animals, e.g., bats, to emit high frequency sounds and then determine the whereabouts of themselves and other objects by hearing the corresponding echoes the sounds produce.
eciliate
Lacking cilia.
ecize
To become established, to spread, to invade. See also: ecesis.
ecocline

The rate of increase or decrease of a variable character in the adaptation of a species, associated with environmental changes. See geocline. 2. The group exhibiting such a gradient. 3. A gradient of ecosystems along an environmental gradient, including both the gradient of natural communities and the complex gradient of environmental conditions. See also: coenocline.
ecologic dominance
The state in communities in which one or more species, by their size, number or coverage, exert considerable influence or control over the associated species.
ecological amplitude (alt. ecological valence)
The variety of environmental conditions within which an organism can survive and replace itself, or a process can function. See also: tolerance, optimum, pessimum.
ecological bonitation
The numerical statement of well-being of an organism or group during a season or in a specific locality. See also: bonitation, biotic potential.
ecological efficiency
A mathematical statement of the ratio between the energy available to an organism or group or group processes, and the energy actually expended. For example, a bear may use less calories to hunt a rabbit during the summer than the calories gained from the rabbit, but winter hunting expends more than is gained, so it is more ecologically efficient for bears to hibernate. A 10 percent gain is average, 20 percent very good, 5 percent typical of the top of the food chain.
ecological equilibrium
See also: the balance of nature, dynamic equilibrium.
ecological equivalence
The case in which two or more species have enough similarities so that any could replace the other in a specified habitat. See also: ecological amplitude, niche.
ecological equivalent
An organism capable of replacing another in a habitat. See also: ecological equivalence, vicariation.
ecological factor
Any variable of the environment that impacts the life of one or more organisms. May be classified into A: climatic, physiographic, edaphic and biotic factors; or B: direct, indirect, and remote factors. See also: biotic, limiting factor, density dependent factor.
ecological longevity
Life span; the average length of life of a species under stated conditions.
ecological pyramid (alt. pyramid of numbers, Eltonian pyramid)
The concept that in most food chains, the number of individuals decreases at each stage, with huge numbers of tiny individuals at the base and a few large individuals at the top, as displayed by millions of plankton, a moderate number of large fish, and a few eagles.
ecology (adj. ecological)
The study of the relation of organisms to their environments.
ecorollate
Without petals.
ecorticate
Lacking a cortex.
ecospecies
A taxonomic species described in terms of its ecological characteristics, usually including two or more ecotypes capable of interbreeding.
ecostate
Lacking a costa.
ecosystem
An interacting complex of a community, consisting of plants and/or animals, and its environmentfunctioning as an ecological unit.
ecotone
A transition area between two adjacent ecological communities containing characteristic species of each, and sometimes, species unique to the area.
ecotourism
Travel undertaken to areas of unique natural or ecologic quality, or the provision of services to facilitate such travel.
ecotype (syn. ecological race, adj. ecotypic)
A subdivision of an ecospecies that survives as a distinct population through environmental selectionand isolation; comparable to a taxonomic subspecies.
ectoparasite
A parasite living on the outside surface of a host, like a tick on a deer.
ectophagous
Refers to an animal that feeds from the outside of a structure, such as a rabbit eating bark from a shrub. See also: entophagous.
ectotrophic
Refers to fungi that grow on the surface covering of roots. See also: endotrophic, mycorrhiza.
edaphic community (alt. edaphic climax)
A climax stage determined by factors related to the soil, pH, drainage, salinity, etc. See also: physiographic climax, biotic climax.
edaphic factor
A condition of the soil that is physical, chemical, or biological that influences organisms growing there. See also: biotic, climatic, ecological factor.
edaphology
The study of soils.
edaphon

An organism that lives in the soil, such as a fungus, nematode, bacterium, etc. 2. The aggregate of organisms in the soil, with the exception of plant parts like roots. See also: plankton.
edentate
With no teeth.
edge
The border between two types of habitat, e.g., between forest and meadow, or stream and prairie, with vegetation of each type.
edge effect
The impact of two diverse communities where they abut, such as where a stream adjoins a prairie. See also: ecotone.
edger
A tool used to trim grass and its roots away from sidewalks and curbs.
EDTA (syn. ethylenediaminetetraacetic acid)
The abbreviation for the white crystalline acid often used as a chelating agent.
eelworms
Microscopic organisms that enter the tissues of plants and cause rotting.
effective soil depth (syn. working depth)
The extent to which roots of plants penetrate readily to reach water and nutrients.
effective temperature range
The spread between the highest and lowest temperatures in which an organism can survive and reproduce. See also: ecological amplitude, tolerance.
effluent
The outflow of water or other fluid. See also: influent.
egg
The nonmotile female sex cell (gamete).
egg-shaped
Shaped like an egg, with the broadest portion below the middle.
eglandular
Without glands.
eilliptic
Widest in the middle and tapering evenly to both ends.
elaiosome
A fleshy, protein-rich “food patch” on some seeds or fruits it is attractive to ants and thus aids dispersal.
elasticity
The capability of a strained body to recover its size and shape after deformation.
elater
A plant structure functioning in the distribution of spores, such bands which contract and expand when moistened or dried to disperse spores and the strap-shaped appendages of Equisetum spores.
electrolyte
A soluble substance capable of conducting ions when dissolved.
elements
Organisms typical of a certain region or habitat, but may occur outside of it, such as a group of prairiespecies found in the eastern part of the united States. See also: ecological factor, vicariation, ecological equivalence.
eligulate
Without strap organs.
ellipsoidal
Widest at or about the middle; margins are symmetrically curved, being narrowed to relatively rounded ends.
elliptic
Narrow at the ends and broad near the center.
elliptical
Oblong with rounded ends.
elongate
Much longer than wide.
eluvial layer
See A horizon.
eluviation
The removal of material from a specified soil horizon by leaching, either in solution or colloidal suspension. See also: illuviation.
emarginate
Having a shallow notch at the extremity.
emasculation

In plants, the removal of male flowers or anthers to prevent self-pollination, such as removing tassels from corn, Zea mays, for hybridization. 2. In animals, castration, the removal of testicles, as done with some bulls to turn them into steers to produce tender meat.
embracing
Clasping at the base.
embryo
The rudimentary plantlet within the seed.
embryo sac
A structure inside the ovule of a flowering plant in which pollination occurs, and in which an embryo begins to develop.
embryogenesis
Formation and subsequent development of plumule, radical, and cotyledons in a plant.
embryoid
A mass of tissue that resembles an embryo.
embryotega
A callus near the hilum of certain seeds, which is detached during germination.
emergence
Refers to outgrowths on the surface of an organ, such as warts, prickles, etc.
emergent (syn. emersed)

Half uncovered. 2. In mosses, of the capsule, when the perichaetial leaves reach but do not overtop it. 3. An aquatic plant with its lower part submerged and its upper part extending above water.
emergent aquatic plants
Plants rooted in shallow water bottoms with the upper leaves and stems above the water surface.
emersed
Raised above the water.
emetics
Herbal medicines that can induce vomiting.
emigrant
One who moves from one area, emigrates, to live in another. See also: immigrant.
emigration
The process of leaving a locality more or less permanently.
emollients
Herbal medicines that can soften skin.
enantiostylous
Refers to flowers whose stamens bend to the left or right, with the styles in the opposite direction.
enation
An outgrowth on the surface of an organ.
encinal
Refers to a grove or forest of evergreen oaks.
enclosure
An area fenced or otherwise encircled to pen in animals, e.g., sheep. An exclosure is similarly treated to shut out other species, like coyotes, so the same area can be both an enclosure and an exclosure.
encystment
A state of inactivity of an organism surrounded by a protective case; metabolism is lowered while resistance is raised to unfavorable environmental conditions. See also: cyst.
end bud
A bud, or sometimes several, located at the end of the twig.
endangered plant
A species adjudged to be threatened with extermination.
endemic
Confined to a small area; limited in geographic distribution.
endemism
The occurrence of highly adapted plants or animals in an area. See also: endemic.
endocarp
The inner layer of a pericarp.
endochory (syn. endozoochory)
Dispersal of an organism, particularly a seed, by an animal which carries it from one place to another, often in its digestive tract.
endogenous
Growing throughout the substance of the stem, instead of by superficial layers.
endoglossum
A crosswise projection from the back wall of a carpel to make it almost completely septate.
endoparasite (alt. endobiophyta)
The existence of a parasite within an organism, like a tapeworm.
endophytic (n. endophyte)

Growing within the tissue of a plant; a plant that grows within another plant, like a fungusendoparasite. 2. A plant that can penetrate a rock, like a lichen.
endoplasma (syn. entoplasm, syn. endosarc)
The inner granular layer of protoplasm in ovum development.
endosperm
The nutritive tissue in a seed of a flowering plant that surrounds the embryo. See also: perisperm.
endospore
An asexual reproductive body found mostly in bacteria.
endostome
The inner row of teeth of a peristome.
endothecium
The inner layers of cells of the capsule.
endotrophic
Refers to fungi that grow within roots. See also: ectotrophic, mycorrhiza.
endozoochore
A propagule like a seed that is dispersed by being carried inside an animal�s body, such as a cherry seed, Prunus, being swallowed by a bird and dropped in feces.
energy flow
The intake, conversion and passage of energy through an organism or ecosystem.
energy transformers
Plants and animals that process and pass on energy, originally transformed from sunlight by plants, from one organism to the next in a food chain. See also: energy flow.
enphytotic
Occurring regularly among the plants of a region, such as an endemic fungus. See also: epiphytotic.
ensiform
Shaped like a sword.
entire
Without toothing or division.
entomophilous
Pollinated by insects.
entophagous
Animals that feed inside of roots, dead leaves, etc. See also: ectophagous.
entropy
The degradation of energy, a measure of the degree of disorder of a system.
envelope (adj. enveloping)
The surrounding part.
environmental clock
The influence of surroundings in initiating activities of organisms or certain processes, such as longer days initiating egg laying in birds.
environmental heterogeneity
The physical or temporal patchiness of the environment, created by disturbances like fire and storms, microclimates, soils, history, and random population variation.
environmental impact
A statement regarding new construction and the consequences it will have on the ecology of the area.
environmental resistance
The limiting influences of environmental factors upon the increase in numbers of individuals in a community. See also: biotic potential, reproductive potential.
Eocene
The second geological epoch in the Cenozoic era–the Tertiary period, that opened about 58 million years ago and lasted for 19 million years.
epappose
With no pappus.
epeirogenesis (alt. epeirogeny, alt. epirogeny)
The deforming of the earth�s crust, producing continents, ocean basins, and great plateaus. See also: diastrophism.
epetiolate
With no petals.
epharmone
An organism which adapted to changes in its environment. See also: ecad, epharmony.
epharmony
The changes of processes or shape of structures by which an organism adapts to an altered environment. See also: adaptation, epharmone, ecad.
ephemeral (syn. transient)

Lasting one day, or more broadly, for a short time. 2. An herbaceous, perennial plant which dies to the ground after flowering and setting seed in spring or early summer.
epibenthic
Living on the surface of bottom sediments in a water body.
epibiotic
An endemic surviving from a former habitat; a relic.
epicalyx
A whorl of bracts outside the calyx.
epicarp
The outer layer of the pericarp or matured ovary.
epichil (alt. epichile, alt. epichilium)
The terminal part of the lip in some orchids, distinctly different from the form of the basal part.
epicole (alt. epibiont)
An organism which attaches itself to a host without either benefit or harm to the host, such as lichenson the bark of trees. See also: commensalism, epiphyte, parasite.
epicormic
Describes new growth (buds, shoots, or flowers) borne on the old wood of trees.
epicortical
Outside the bark.
epicotyl
The stem of a seedling between the cotyledons and the first true leaves.
epidemic
The pervasive spread of organisms which are parasitic, predatory, or damaging.
epidermis
The outermost layer of cells covering the plant.
epigeal
Of or relating to the emergence of cotyledons above the surface of the ground. See also: hypogeal.
epilia
A population of air plants, epiphytes. See also: epicole.
epilimnion
The upper layer of a lake which is disturbed by winds, lying above the thermocline. See also: hypolimnion.
epilithic
Growing on rocks.
epinasty (adj. epinastic)
In plant physiology, the state in which more vigorous growth occurs in the upper surface of an organ, such as in an unfolding leaf, causing a downward curvature. See also: hyponasty.
epiorganism (alt. supraorganism)
A natural group made up of similar individuals, like a hive of bees or a stand of maples, Acer. See also: community, population.
epipetalous
Borne on or attached to the petals.
epipetric
Growing on rock.
epiphragm
A membrane covering the mouth of the deoperculate capsule.
epiphyll
An organism that grows on leaves. See also: epiphyte.
epiphyllous
Growing on a leaf, usually through vegetative reproduction.
epiphyte (adj. epiphytic, syn. aerophyte)
A plant growing attached to another plant, but not parasitic; an air plant.
epiphyton
A collection of organisms scattered over surfaces submerged in water, that may later become mechanically associated. See also: lasion, periphyton.
epiphytotic
Common, even epidemic, among plants in general, as some fungus diseases like mildew. See also: enphytotic.
episepalous
Borne on or attached to the sepals.
epispastics
Herbal medicines which can cause blisters.
epistasis
Suppression of the effect of a gene by another, nonallelic gene.
epithalassa
The upper layer of water above the thermocline, where stratification occurs because of water temperature in the ocean.
epithelium
Cellular tissue which covers a surface or lines a tube to assist secretion production and help assimilate nutrients.
epithet
The part of a scientific name designating a species or lower division of a genus. For example, in Cyranthus mackenii var. cooperi, mackenii is the species epithet and cooperi is the variety epithet.
epizoan
An epiphyte animal.
epruinose
Lacking bloom on the surface.
epsom salts
Bitter crystalline hydrated magnesium sulfate, sometimes used as a soil amendment.
equal
Of the same length, as in ‘sepals equal petals.’ It means that two plants are the same length–not that they are indistinguishable, as when two plants are so inseparable they are called tepals.
equidistant
The same measurement apart.
equilateral
Equal on both sides of an axis.
equitant
Astride; refers to conduplicate leaves which enfold each other in two ranks, alternating one above the other on opposite sides of the stem, as occurs on an iris.
eradicate (n. eradication, syn. exterminate)
To remove entirely; to pull up by the roots.
erecto-patent
Midway between erect and patent.
erect
Vertical or upright
eremean
Belonging to regions of low, irregular rainfall.
erianthous
Having woolly flowers.
ericaceous
Refers to plants of the family Ericaceae that require an acid soil, generally with a pH of 6 or less.
ericoid
Like a heath.
erinous
Prickly, with sharp points.
erose
Irregularly cut away as if gnawed.
erosion (v. erode)
Slow destruction of soil or rock by the action of water, wind, or ice.
errhines
Herbal medicines applied in the nose to promote discharge of mucus.
erumpent
Appearing to be ready to break through.
escape
An exotic plant that has spread from cultivation and grows successfully in the wild.
escarpment (alt. scarp)
A long inland cliff or steep slope formed by erosion, often by waves, or sometimes by a fault.
esorediate
Lacking soredia.
espalier
A plant trained to grow flat against a wall or framework. To train a tree or shrub with its branches growing in a two-dimensional plane, flat against a building or fence.
essential oil
Any volatile plant oil used in perfume or flavorings.
estipulate (alt. exstipulate)
With no bract at the base of the petiole.
estivation
Stagnating or otherwise nonfunctional during the summer months. See also: hibernation.
estrogenic
Herbal medicine that stimulates female hormone production or replaces it with plant hormones.
ethnobotany
The plant lore and agricultural customs of a people; the systematic study of such lore.
ethnopharmacology
The study and practice of medicines used by a people.
ethylene
A colorless flammable hydrocarbon gas occurring in plants, acting as growth regulator and fruit ripener.
etiolate (n. etiolation)
To alter the natural development of a plant by excluding sunlight, often resulting in pale or bleached foliage. Etiolation is one method of layering for propagation; the shaded parts turn white or cream and develop no leaves.
eucamptodromous
Describes leaves having pinnate venation in which the secondary veins do not terminate at the marginsbut which gradually diminish inside the margin, connected to the superadjacent secondary veins by a series of cross-veins without forming prominent marginal loops. See also: acrodromous, brochidodromous, semicraspedodromous.
Euglenophyta
A taxonomic division containing euglenoids and Euglena. All are single-celled and found mostly in fresh water. They can contract the vacuole and have a single flagellum.
eukaryote
An organism composed of one or more cells containing visibly evident nuclei and organelles, including all organisms except viruses, bacteria, and cyanobacteria. See also: prokaryote.
eusporangiate
Refers to ferns having sporangia with walls thicker than one cell. See also: leptosporangiate.
eusporangium
A thick-walled sporangium originating from several epidermal cells.
eutrophic
Rich in dissolved nutrients, photosynthetically productive and often deficient in oxygen during warm weather.
evanescent
Disappearing at maturity.
even-pinnate
A pinnately-compound leaf lacking a terminal leaflet.
everlastings
Plants with flowers which hold their color and shape when dried.
evolute
Opened, unfolded.
evolution
Organic evolution is any genetic difference in organisms from generation to generation.
ex situ conservation
A conservation method that entails the removal of seed, pollen, sperm or individual organisms from their original habitat, keeping these resources of biodiversity alive outside of their natural environment.
exalate
Lacking an ala.
exalbuminous
Without albumen.
exappendiculate
With no appendages.
exarate
With grooves.
exarillate
Without an aril.
exceed (alt. exceeding)
Refers to parts being compared in size where one is longer than the other.
excentric
Off center.
excrescence
Abnormal development, an outgrowth.
excurrent

A growth habit with a single vertical trunk, the branches obviously secondary. 2. Describes a costarunning out beyond the lamina of a leaf. 3. Running out, as a nerve of a leaf projecting beyond the margin.
excurved (n. excurvature)
Curved out and away from the central part.
exedra

An open or colonnaded recess used for conversation, often semicircular and furnished with seats or a long bench. 2. A semicircular bench with a solid back.
exfoliate
To peel off or shed, as the thin layers of bark.
exfoliating
Cleaving off in thin layers.
exiguous
Small and narrow.
exindusiate
Lacking indusia.
exine
The outer layer of the wall of a pollen grain or spore.
exocarp
The outer layer or skin of a pericarp.
exogenous

Grown or made outside the body. 2. Growing by annular layers near the surface, as bark. 3. Produced by growth from superficial tissue.
exostome
The outer row of teeth of a peristotne.
exotic
Not native, from another area.
exotic species (syn. alien species)
A species that is not native to a particular geographic location, but may have become naturalized there.
expanded
Increased in area or size.
expectorants
Herbal medicines which can loosen mucus from the throat.
explanate
Flat, fanned out.
exserted

Projecting beyond an envelope, as stamens from a corolla. 2. In mosses, elevated above the surrounding parts of the capsule, when the perichaetial leaves do not reach as high as their base.
exsiccated
Dried.
exstipitate
Lacking a stipe.
exstipulate
Having no stipules.
extant
Still in existence; not extinct.
extensile
Capable of enlargement or extension.
extinct
No longer living; no longer in existence.
extinction (syn. extirpation)
The evolutionary end of a species or the loss of a species in a large area such as a state or country, caused by the failure to reproduce and the death of remaining members of the species.
extirpated
Literally, plucked up by the roots, exterminated. Often used to indicate that a species once lived in an area, and no longer does, usually because of changes in habitat.
extirpation
Wiped out, completely destroyed. Carries the connotation of differing from extinction in that it is more often due to human causes.
extra-axillary
Arising on the side of a stem outside of or opposite to an axil.
extract
A product prepared by removing essential constituents of a plant, such as oils for flavorings.
extrafloral
Beyond the essential parts of the flower, as nectaries, bracts, etc.
extrastaminal
Outside the stamens.
extratropical
Outside the tropics.
extravaginal
Describes a shoot which arises from an axillary bud and breaks through the sheath of the subtending leaf.
extrorse
Facing outward.
exudate (syn. sweat)
Matter oozing from a surface.
exvaginate
Outside the sheath.
eye

The center of a flower, if a different color from the rest. 2. A stem cutting with a single bud. 3. The undeveloped bud on a tuber.
eye-spot
A red cytoplasmic structure sensitive to light.
F layer
Sometimes used to designate the upper layer of soil litter whose origin and age can still be determined, although it is partially decomposed. See also: A horizon, H layer, L layer.
F1 hybrid
First-generation cross between two pure-bred strains. These hybrids produce seeds that are not true to type. See also: filial generation.
F2 hybrid
Second-generation cross between two F1 hybrids. See also: filial generation.
faciation
A part of a climax association which lacks some of the dominants of the normal association due to slight differences in environmental fundamentals.
facies

A variation of the biotope, differing from the typical conditions in minor ways. 2. A deviation in a community like a dogwood, Cornus, or shrub facies in an oak forest. 3. The top to bottom appearance of a plant, including foliage, flowers, fruit, roots, etc. See also: faciation.
facultative
Capable of functioning under varying environmental conditions, e.g., a parasite which can survive with or without a host. See also: obligate.
facultative apomict
A plant that can reproduce either sexually or asexually (apomixis.)
fairy ring
A circle of mushrooms or toadstools growing from underground mycelium, which is usually surrounded by a ring of vegetation and is rather common in grasslands and sometimes in forests.
falcate
Scythe-shaped; curved and flat, tapering gradually.
fall line
A line connecting the points where rivers leave the uplands as they flow to the lowlands, indicated by sharper slopes and waterfalls.
fallout
The dropout from the air of solid materials, particularly in reference to radioactive dust from nuclear explosions.
fallow
Refers to cropland left standing without cultivation except to destroy weeds and accumulate water and nutrients for a later crop.
falls
Pendulous outer petals, as those found on an iris.
false annual ring
An extra growth ring produced in a season with a drought and then more rain.
family
A group of plants sharing common features and distinctive characteristics and comprising related genera; the taxonomic category above genus and below order.
fan-compound
A compound leaf with leaflets radiating from a single point; palmate-compound.
fan-lobed
Major lobes radiating from a single point; palmate-lobed.
fan-veined
Main veins radiating from a single point; palmate-veined.
fancy
Term used by the International Code of nomenclature for cultivated Plants. It is applied to cultivars, grexes, and hybrids, as opposed to natural species.
fangy
Describes forked roots.
farina
A waxy-appearing substance exuded from glands.
farinaceous (syn. farinose)
Containing starch; powdery, starch-like.
farinose
Covered with a waxy, whitish powder; bearing farina.
fasciation
An abnormal flattening or coalescence of stems or leaf stalks.
fascicle
A close bundle or cluster.
fasciculate
In close bundles or clusters.
fasciculated root
A fibrous root where some of the branches are thickened.
fastigiate
Describes branches that are erect and near together, forming a columnar shape.
father plant
The species from which pollen was obtained to create a hybrid.
fauces (sing. faux)
The throat of a flower in which the petals are united at least at the base, especially when the throat is distinguished in some way such as color, scales, etc.
fauna
A collective term, including all the kids of animals in an area or geologic period.
faunal region
An area with characteristic kinds of animals.
faveolate (alt. favose)
Honey-combed.
favulariate
Finely ribbed, the ribs separated by zig-zag furrows.
feather-compound
Midribs of main leaflets branching from a central midrib at several points in a feather-like pattern; pinnate-compound.
feather-lobed
The main lobes more or less at right angles to the midrib, not radiating from a central point; pinnate-lobed.
feather-veined
Describes leaves whose veins all arise pinnately from a single midrib.
feces
Animal droppings, manure, scats.
fecundity
Productivity of an organism regarding fertility structures, like eggs, sperm, pollen, etc.
feedback
Any partial reversion of the effects of a given process to its source, such as leaves falling to the ground and furnishing calcium for uptake by the roots of the plant.
feeder root
One of the numerous small roots of a plant, through which moisture and nutrients are absorbed from the soil.
felted-tomentose
Woolly and matted, the hairs curling and tangling tightly to the surface.
fen
Low marshy ground containing peat that is relatively rich in mineral salts and is alkaline rather than acidic. It is usually found in the upper parts of old estuaries or around fresh water lakes, with vegetation quite different from that of moors.
fenestrate
Having openings, perforations or translucent areas.
Feng Shui
An ancient Chinese discipline combining town planning, environmental impact, architecture and interior decoration. Time, space, and action are designated to increase energy, harmony, healing, etc.
feral
Refers to an organism that escapes from cultivation or domestication and becomes naturalized.
fermentation
The chemical change of organic substances by organisms, such as yeast turning sugar into alcohol and carbon dioxide.
fern ally (pl. fern allies)
A nonflowering, vascular plant belonging to one of the following classes: Equisetopsida (horsetails) Lycopdiopsida (club mosses, spike mosses, and quillworts) and Rhyniopsida (whisk fern).
ferruginous
Rust-color.
fertile
Capable of producing fruit; or productive, as a flower having a pistil or an anther having pollen.
fertility

The reproductive capacity of an organism. See also: fecundity. 2. The property of soil that allows it to provide available nutrients and moisture to plants.
fertilization (alt. fertilisation)
The union of male and female gametes.
fertilizer
Synthetic or natural organic substances providing essential plant foods. It most often includes nitrogen (N), phosphorus (phosphate; P), and potassium (potash; K).
fetid (alt. foetid)
Carrying an unpleasant odor.
fibril
A short simple lateral branch of a lichen.
fibrillate
Finely fibrous or with hair-like lines.
fibrillose

Furnished or abounding with fine fibers. 2. Having fibrils.
fibrous
Composed of or resembling fibers. Fibrous tissue: a tissue formed of elongated thick-walled cells.
fibrous root
A root that has no prominent central axis and that branches in all directions.
fibrovascular
Composed of woody fibers and ducts.
fiddlehead
One of the young unfurling fronds of some ferns that are often eaten as greens.
fidelity
The regularity with which a species occurs in certain plant communities, expressed in a five-part scaleranging from the most to least common: 5) exclusive, 4) selective, 3) preferential, 2) companion or indifferent, 1) accidental or stranger.
field border plantings
Vegetation established on the margins of fields to conserve soil and to provide food and shelter for wildlife, like native shrubs. See also: filter strip.
field crops (syn. cash crops)
Agricultural production such as grains, hay, root crops and fiber in contrast to vegetables and fruits. See also: truck crops.
field equipment
Gear necessary to collect botanical specimens in the wild, such as maps, diggers and clippers, trowels, hand lens, collection bottles and bags, insect repellents, cameras, etc.
field expedition (syn. field trip)
A journey planned to collect flora of a specific area or habitat.
field layer
The low stratum of plants like grasses, forbs and dwarf shrubs. See also: canopy layer.
field preservatives
Materials used to hold wet plant samples for long periods of time, or to hold plant specimens in tropicalareas. Examples are 2:3 commercial 40 percent formaldehyde and water; 1:2 formaldehyde and 70 percent alcohol; 40-50 percent alcohol; 1-2 percent aqueous solution of oxyquinoline sulfate. Specimens may be dipped, sprayed, or brushed with one of these, then enclosed in an airtight container.
field records (syn. field notes, syn. field data)
A pocket-sized notebook which holds complete data on a field trip, including the precise location of places explored, dates, flora collected with any assigned numbers such as a tube of pollen or seeds, information on such things as a location in shade or sun, insects found on plants, moisture, bloom or fruit and their colors, and nearby plants, etc.
field strip cropping
A specialized strip cropping where crops are planted in parallel bands across a slope but do not follow contour lines; bands of grass or other close-growing species are alternated with the bands of cultivatedcrops.
field test (syn. field laboratory)
An experiment conducted under regular field conditions, less subject to control than a precise contained experiment.
filamentous

Made up of filaments or bearing them. 2. A form of lichen containing filamentous algae and fungi.
filial generation
In hybrid offspring, the first cross is designated as F1; the second cross as F2, etc.
filiform (syn. thread-like)
Thread-shaped; long, slender, and terete.
filmy fern
A fern of shady places with exceptionally thin frond leaflets, usually of the family Hymenophyllaceae.
filter bridge
A land connection that remains in place for a demonstrated period of time, restricted in extent, limiting the kinds of organisms which can migrate over it, such as the Bering Strait in the Pleistocene period. See also: corridor, sweepstakes bridge.
filter strip (syn. greenway, syn. greenbelt, syn. beltway, syn. buffer strip, syn. buffer zone, syn. vegetated filter zone)
A permanent strip of low plantings on the lower side of a field that is sufficiently wide and dense to slow the movement of runoff, allowing deposition of silt on the strip rather than its transport to a stream or pond. See also: field border plantings.
fimbriate (alt. fimbriated, n. fimbriation)
Fringed; having the edge or extremity bordered by slender processes.
fimbrillate
Having a minute fringe.
fine-textured soil
A soil that consists mostly of silt and clay, with little or no sand or gravel.
fines
Materials from rock and organic extraction that are often carried away by the action of wind or water.
fire control line
A designated margin along which a wildfire, if possible, is not allowed to pass; often backfires are started here.
fire hazard
The risk level of a fire starting under prevailing climatic conditions, often simply low, average, or high.
fire towers
Structures in forests which rise above the tree tops and are staffed during fire danger periods.
fireblight
A bacterial disease of rose family.
firebreak
A strip of land where flammable materials have been removed to check or stop a creeping or running wildfire.
firn
Granular snow, usually compacted; a ski term for last year’s snow. See also: neve.
first bottom
The flood plain that is closest to a stream or is at the lowest elevation above a stream; the first to be flooded.
firth
A term commonly used in Scotland that means a long narrow arm of the sea or the mouth of an estuary.
fishway (syn. fish ladder)
A sloping structure over which water is allowed to flow, to help fish in breeding season to ascend a stream which forms a waterfall.
fissile
Easy to split.
fissured
Having deep cracks.
fistular
Hollow and cylindrical.
fitness
The level of adaptation of an organism to its environment.
fitness of environment
The suitability of a habitat for preserving life.
flabellate
Fan-shaped.
flachmoor
Wasteland with a flat to slightly depressed surface, with peaty soils that are poor in salts and have an acidic reaction.
flagellates
Cells with flagella but no cell wall.
flagelliflorous
Bearing flowers at the ends of long, pendulous branches.
flagelliform
Whip-like in shape.
flagellum (pl. flagella)
Tiny whip-like appendage that is capable of movement.
flaky
Bark with loose scales which are easily rubbed off.
flank
The side of anything, like the west flank of a mountain.
flats
Low, exposed land of a lake delta or a lake bottom; or often brushy land along the floodplain of a stream.
fleshy
Succulent or juicy.
flexible
Easily bent or twisted.
flexuose (alt. flexuous)
Zigzag; bending from side to side; wavy; sinuous.
floating-leaved plants
A free-floating plant such as duckweed, Lemna; or one with roots in the water bottom but having leaves that float on the surface like the water lily, Nymphaea.
floccose
Clothed with locks or flocks of soft hair or wool.
flocculate (n. flocculation)

To cause to form lumps or masses, as in soil. 2. The rapid precipitation of large amounts of a solute out of a solvent.
flocculation
The coming together of suspended colloidal material or very fine particles like silt, into larger masses called floccules.
flocculent
Resembling wool especially in loose fluffy organization.
floodplain (alt. flood plain)
The almost level land forming the floor on either side of a stream in a valley, often subject to flooding.
flora
All plants as a group; the particular plants of a region.
floral
Belonging to or associated with a flower.
floral cup
A cup-shaped or tubular part of a flower.
floral envelope
The perianth.
floral region (alt. floristic region, alt. floristic area)
An area with a degree of homogeneity of species existing because of similarities in topography, climate, etc., in the region.
floral shovel
A round-pointed shovel.
floral snip
Scissors specialized for delicate jobs, such as harvesting herbaceous flowers.
floral tube
A tube formed in some flowers by the fusion of the perianth and the stamens.
floret
A small flower, usually one of a dense cluster. See also: flowerhead.
floricane
A biennial stem typical of Rubus, which grows the first year, and bears fruit and dies the second year.
floriculture
The cultivation of ornamental plants, especially flowering plants.
floriferous
Flower-bearing; blooming profusely.
florigen
A hormone which appears to be produced in the leaves, then translocated to apical meristems where it initiates the formation of flowers.
florist
One who sells flowers retail, usually obtaining the blossoms from a middle man.
floristic composition
The array of plant species that occurs in a community or region.
floristic element
Species that are characteristic of a designated region but may also occur in a different one, e.g., a midwest united States species growing in the upper levels of the Smoky Mountains is a midwest element of those mountains.
floristic study
A complete listing of the plants from a given jurisdiction, e.g., a state, or from habitats such as lake shores or marshes, or a listing of all of the species within a genus or other delineation.
floristic territory
A geographic area characterized by the common occurrence of a number of more or less endemicspecies, although other species may be present.
flower

The reproductive structure of a flowering plant consisting of a pistil and/or stamen, and usually including petals and sepals. 2. Often applies to the reproductive organs of mosses.
flower bed
An area planted with flowering plants and surrounded by lawn, structures, etc.
flower box (alt. window box)
A container attached to a porch railing or window sill planted with colorful species.
flower bud
The bud which contains a flower or a cluster of flowers.
flower head
A group of florets.
flower induction
The initiation of the production of flowers, possibly stimulated by florigen.
flowering dates
The period(s) of the year when a species can be expected to produce blossoms.
fluctuation
A relatively atypical departure from more average conditions, but still within normal limits.
flume

In the united States, a ravine or gorge with a stream running through it. 2. A man-made open conduit, often made of wood or concrete, for the purposes of irrigation, power, etc. It operates by transporting liquids across a stream or depression which intersects the course of the conduit.
flush cut
A pruning cut to remove a tree limb in which the cut is completely flush with the tree. The resulting scar is too large to heal efficiently.
fluted
With rounded, longitudinal grooves or ridges.
flyway
A migratory pathway of birds, like the Mississippi flyway following the Mississippi River.
foehn
A Swiss term for a wind blowing down a mountainside, especially if it is warm for the season. See also: chinook.
fogger
A device used to apply a pesticide in suspension in water or air.
foliaceous
Leaf-like in texture or appearance.
foliage
The leaves of a plant taken collectively.
foliar diagnosis
Evaluation of the nutrients in a plant, or the plant nutrient requirements of a soil, by analyzing the leaves.
foliar feeding (alt. foliar fertilizing)
The process whereby plants are fertilized by application of liquid onto the leaves rather than through the soil.
foliate
Bearing leaves.
folic acid (syn. pteroylglutamic acid)
A member of the B vitamin complex, found mostly in the leaves of plants.
foliolate
Bearing leaflets.
follicular
Like a follicle.
food chain
An ever increasing progression of edibles, from microscopic plankton which are eaten by small animals, which in turn are eaten by larger animals, and so on, up to the largest predator. The animals all die and are digested by fungi and bacteria, starting the chain all over again.
food cycle (alt. food web)
All the interconnecting food chains in a community.
food niche
The location of a particular organism in a food cycle.
food pyramid
A theoretical graphic illustration showing the immense number of producer organisms at the base and the progressively decreasing numbers of herbivores and carnivores toward the peak.
forage acre
A theoretical concept demonstrating the quantity of feed on an acre of land which is totally coveredwith vegetation and completely utilized under proper management.
forage acre requirement
The number of forage acres needed for the maintenance of X numbers of mature grazing animals for a specified period of time.
forage ratio
A mathematical statement regarding the percentage of a prey species present in the food of a predator species, which is divided by the percentage of the prey species present in the habitat.
forage volume

The total parts of plants that can be reached by grazing or browsing animals. 2. A measure of the yield of feed, the total amount produced on a certain range during a year.
foraminifers
Small marine animals with calcareous shells which are perforated with minute holes for extension of pseudopods.
forb
A nonwoody plant other than grass, sedge, or rush. See also: herb.
foredune
A low pile of sand, often held in place by dune grass, bordering the shore of a sandy lake or sea.
forensic
Relating to the application of scientific knowledge to legal problems, as a forensic entomologist helps place time of death by developmental stages of insects on the deceased.
foreshore
That portion of the beach occupied on a daily basis by tides.
forest cover
The living plants and dead organic matter on the floor of a forest, sometimes restricted to only the plants. See also: basal cover, duff.
forest edge
The boundary or ecotone of a forest where it meets with some other kind of vegetation such as a marsh.
forest floor
The deposits of plant material such as dead leaves and branches on the ground in a woodland. See also: duff, forest cover.
forest fragmentation
The continuing splitting of large forested areas by highways and residences, which changes the habitat. It affects the acclimation and persistence of wildlife and indigenous plant species.
forest influences
A total of the effects or reactions of a woodland on the environmental conditions, such as providing duff as compost, and maintaining uniform stream flow.
forest type
A stand of trees that is fundamentally similar throughout and can be predicted to occur elsewhere under comparable conditions. It includes temporary, permanent, climax, and cover types.
forked
Divided into nearly equal branches.
forma (abr. f.)
The rank of taxa below variety; the narrowest taxon; a plant which retains most of the characteristics of the species, but differs in some way such as flower or leaf color, size of mature plant, etc. A forma is added to the specific binomial and preceded by “f.”, such as rubra in the epithet Cornus florida f. rubra.
form
A small but constant variation within a population of plants, such as a white-flowered plant in a normally purple-flowered population.
formenkreis
A series of related forms pinpointed geographically that originated entirely or primarily by geographic isolation. See also: speciation, evolution.
fortnight
Two weeks, fourteen nights.
fossorial
Refers to animals that burrow in the soil, e.g., voles.
foundation planting
Those shrubs and flowers planted close to the house which make a smooth transition to the surrounding landscape.
founder principle
The theory that an isolated population, perhaps on an island or cut off in other ways, holds a small number of settlers with a limited gene pool, which is the first step toward new species.
foveolate (alt. foveate)
Honeycombed; pitted.
fox fire
Phosphorescent light caused by a fungus on decaying wood.
fragment
Refers to communities, a stand so small that it lacks sufficient species composition and other characteristics associated with a community. See also: population.
fragmented structure
Refers to a soil composed mostly of particles that exhibit well defined faces and edges. See also: granular structure.
fragrance garden
A garden consisting of flowering plants and herbs noted for pleasant odors. Often used in recreational/educational areas for the visually impaired, and for residential yards used at night.
frass
Debris and fecal matter produced by insects.
free-central
Describes placentation where the ovules are borne on a free-standing central placenta within the ovary.
French drain
A channel leading from an overly wet area or to a dry area to transport water as needed. The deep narrow channel is filled with stone to allow water to pass, but to also provide a firm surface.
frequency (alt. frequence)

The level of regularity found in the distribution of individuals of a species in an area, especially in a stand. See also: constancy, stand. 2. A general term for a group of plants with more or less regularity of physiognomy, composition, and habitat. See also: community, association.
frequency class
One of the small groups into which a frequency index of the various species in a stand may be classified.
frequency index
A mathematical expression using the percentage of frequency; e.g., a species inhabiting 16 of 20 sample areas has a frequency index of 80 percent.
friability
Desirable texture of soil that allows it to crumble into small particles rather than large clods.
frigid zone
That part of the earth lying north of the arctic Circle or south of the Antarctic Circle. See also: temperate zone, tropic zone.
frigorideserta
Tundra; cold arctic or antarctic areas, and alpine communities.
fringed
Fimbriate.
fringing forest
A strip of woodland along a stream or body of water. See also: gallery forest.
frondose
Leafy, frond-like.
front
The border between warm and cold air masses at the earth’s surface.
frost

A covering of minute ice crystals on a cold surface. 2. The temperature low enough for this to occur. See also: hard frost, light frost.
frost resistance
The capacity for survival of a plant when ice crystals form in the cells. See also: hardening.
frostless season (alt. frost free days)
The time between the last spring frost and the first autumn frost.
fructiferous
Capable of bearing fruit.
fructification
The act or organs of fruiting.
frugivore
An animal that eats fruit.
fruit

The mature ovary of a seed plant. 2. Sporangia; often applied to the sporophyte.
fruit harvester
A basket made of wire with curved tips, fastened to a long handle, to catch fruit and pluck it from the ground.
fruit-dot
A sorus.
fruiting spur
On a fruit tree, a short twisted branch with rings around it, which flowers and produces fruit.
frutescent (alt. fruticose)
Resembling a shrub.
fruticeta
Vegetation type made up of scrubby forest.
fruticose
Shrub-like, bushy, with many stems rather than a single trunk; a form of lichen which appears shrubby or hair-like.
fruticose lichen
A lichen with a thallus more than 10 cm. (4 inches) tall, like Cladonia rangiferina, reindeer moss.
fucoid
Resembling a seaweed, especially Fucus.
fugacious
Falling or fading very early; short-lived.
fully double
Flower heads with multiple rows of ray florets; the disc florets are immature and completely covered by the central rays when the flower is at its prime stage.
fully revolute
Rolled backwards with margins touching or overlaping.
fully stocked
Refers to a stand that contains as many trees or other materials of the species and ages as the site can support. See also: overstocked, understocked, carrying capacity.
fumarole
A hole in the earth from which heat and gasses escape under pressure.
fungicide (adj. fungicidal)
An agent to inhibit the growth of, or destroy fungus.
fungivorous
Refers to organisms that eat fungi.
fungoid
Resembling a fungus, mushroom-shaped.
fungus
A nonflowering plant of the kingdom Fungi, all lacking chlorophyll.
funicle (alt. funiculus)
The free stalk of an ovule or seed.
funiculus
The stalk of the ovary in plants.
furfuraceous
Covered with bran-like scales.
furrow dam
A small earth blockade for holding water within a furrow. See also: lister.
furrowed
With channels running lengthwise, including both sulcate grooves and striate thread marks.
fusarium (pl. fusaria)
Any of the pathogenic, imperfect fungi of the genus Fusarium which infect both plants and animals.
fuscous
Dusky, grayish-brown.
fusiform
Spindle-shaped; swollen in the middle and narrowing toward each end.
fusiform root
A taproot which tapers at the top as well as the bottom, like a white radish.
fusoid
Somewhat spindle-shaped.
fynbos
A South African term for sclerophyll vegetation on plateaus and mountains, similar to macchia of the Mediterranean region and chaparral of California.
galactosidase
An enzyme which cuts the glycosidic bond between the sugar galactose and molecule. There are two types, the alpha galactosidases and the beta galactosidases.
galea
A hooded or helmet-shaped portion of a perianth, as the upper sepal of Aconitum, and the upper lip of some bilabiate corollas.
galeate
Helmet-shaped; having a galea.
gall
An abnormal growth or swelling caused by insects, fungus, etc.
gallery forest
A narrow woodland developed along a stream because of constant and sufficient moisture through a grassland or other open vegetation.
galls
Corky swellings.
gametangium
The organ producing the gametes.
gametophyte (syn. gametophore)
That part of the plant which bears the gametes or sexual cells. In mosses, all of the plant except the “fruit,” or seta and capsule.
gamopetalous
Having the petals of the corolla more or less united. See also: polypetalous.
gamophyllous
Composed of coalescent leaves or leaf-like organs.
gamosepalous
Having the sepals united.
garden
The proper term for landscaped area or yard. To many Americans, the term refers only to the vegetable garden.
garden hoe
A lightweight hoe for weeding, etc., in home or truck gardens.
garrigue (syn. phrygana, syn. batha)
Stony or sandy–often over-grazed–hillsides, similar to maquis, but hotter and drier still.
gazebo
A free-standing roofed structure usually open on the sides.
gelatinous
Having the consistency of jelly or gelatin.
geminate
Equal, in pairs.
gemmiferous (syn. gemmiparous)
Bearing gemmae.
gemmiparous
With vegetative buds.
gene bank
A facility existing for the ex situ conservation of seeds, tissues, or reproductive cells.
genetic diversity
Variability in the genetic makeup of individuals within or among species; the hereditary variation within and among populations.
genetic dominance
The influence exerted by a dominant gene or allele, like wrinkled seeds being dominant over smooth.
genetic drift
Random changes in isolated populations, or in the frequencies of certain genes, which cannot be accredited to selection, mutation, or migration. See also: natural selection.
geniculate (alt. genticulate)
Bent abruptly, like a knee.
geniculum
A sharply bent joint or node.
genotype
The genetic constitution of an organism, acquired from its parents and available for transmission to its offspring. See also: phenotype.
genus (pl. genera)
A group of closely related species. The taxonomic category ranking above a species and below a family.
geophyte
A plant whose perennating buds are found underground, usually attached to a bulb, corm, tuber, etc.
geotropism
An organism’s growth determined by the force of gravity, as roots growing downward and stemsupward.
germ cell
A reproductive cell, or gamete, or a cell that will develop into a reproductive cell, such as a spermatocyte or an oocyte.
germ plasm
The cytoplasm of the germ cell, particularly that containing the genetic material.
germination
The sprouting of a plant seed.
germination rate
The proportion of seeds in a given seed lot that are likely to germinate under favorable conditions.
germplasm
The genetic material with its specific molecular and chemical makeup that comprises the physical foundation of the hereditary qualities of an organism.
gesneriad
Any of the mostly tropical plants of the family Gesneriaceae, including gloxina and African violets.
gibberellic acid
A hormone used to promote plant growth, especially that of seedlings, and obtained from the fungusGibberella fujikoroi.
gibberellin
Any of several growth-regulating plant hormones produced by seeds, mosses, ferns, algae, and fungi.
gibbosity
A swelling of moderate extent and asymmetrical character, chiefly at or near the base of an organ.
gibbous
Protuberant or swollen on one side.
girdle

To intentionally remove a circle of bark and cambium from a tree in order to kill it. 2. To encircle tightly, eventually choking of nutrients and causing the death of a plant or limb.
girdling root
A root that has become wrapped around the trunk of the plant which inhibits the uptake of nutrients; usually occurring in plants grown in containers.
glabrate
Somewhat glabrous, or becoming glabrous.
glabrescent
Becoming glabrous in age.
glabrous
Smooth; not rough, pubescent, or hairy.
glade
An open grassy area surrounded by woods.
gland-dot
A tiny gland or pore, usually secreting fluid.
glandular
Bearing glands or of the nature of a gland.
glandular-toothed
Teeth that bear glands.
glaucescent
Somewhat glaucous, lightly coated with a fine bloom.
glauconite
A greenish mineral consisting of a silicate of iron, magnesium, aluminum or potassium found in greensand and often used as a fertilizer or soil supplement.
glaucous
Bluish white; covered or whitened with a very fine, powdery substance.
globose
Globe-shaped.
globular
Spherical or nearly so.
glochid
A barbed hair or bristle.
glomerate
Compactly clustered.
glomerule (adj. glomerulate)
A small compact cluster.
glucose
A monosaccharide sugar widely found in plant and animal tissue.
glumaceous
Furnished with or resembling glumes.
glume
A chaff-like bract; specifically one of the two empty chaffy bracts at the base of the spikelet in the grasses.
glutinose (alt. glutinous)
Viscid, sticky.
glycosidic bond
A bond between a sugar and another organic molecule by way of an intervening nitrogen or oxygen atom.
Golgi complex (syn. Golgi apparatus, syn. Golgi body)
A cytoplasmic organelle that consists of a stack of smooth membranous saccules and associated vesicles and that is active in the modification and transport of protein.
grade adjustment
Changing existing levels of ground areas.
grader (alt. roadgrader)
A truck which smooths the surface of gravel roads and raw soil.
graft
To insert a section of one plant, usually a shoot, into another so that they grow together into a singleplant; the plant formed from grafting.
graft chimaera (alt. graft hybrid)
The mingling of tissues belonging to the stock and scion through grafting in a nonsexual manner. These are designated in formulae by a crucifix type cross which appears as ““, e.g., _Laburnocytisus adamii (Laburnum anagyroides Chamaecytisus purpureus).
grafting
Method of propagation for trees and shrubs by inserting a section of one plant, usually a shoot, into another so that they grow together into a single plant.
graminaceous
Related to grain-bearing plants.
granular structure
Refers to a soil composed mostly of particles that exhibit rather indistinct faces and edges. See also: fragmented structure.
granulose
Composed of or appearing as if covered by minute grains.
granum (pl. grana)
Structures in chloroplast that consist of thylakoids.
grass catcher
A container fitted onto a lawn mower to catch grass clippings while mowing.
gravitational water
Water held in large pores in soil that drains away or leaches when underdrainage is free.
gray mold
A fungus disease characterized by the gray hairy appearance of affected parts.
graze

To feed on grasses and forbes. 2. Grasses and forbes.
green drought
A weather term describing a period of enough rain to keep shallow-rooted plants alive, although the water table continues to recede.
green manure
The plowing under of living plants so that their decay will increase the humus of the soil.
greenhouse
An enclosed structure, usually made of glass (to trap solar radiation), that allows for the regulation of temperature and humidity and is used to propagate and grow plants.
greensand
A sand or sediment that consists largely of dark greenish grains of glauconite, usually mixed with clay or sand.
gregarious
Growing near together or clustered, but not in close tufts or mats.
grex
A group name for all plants derived from crossing the same two or more parent species; the herd or hybrid swarm.
grooved
A general term for sulcate or striate.
grotto
An artificial recess or structure made to resemble a natural cave, often having running water.
ground layering
Another name for tip layering.
groundcover
A plant with a low-growing, spreading habit, grown specifically to cover the ground.
groundwater
Subsurface body of water body in the zone of saturation; That portion of the water beneath the surface of the earth that can be collected with wells, tunnels, or drainage galleries, or that flows naturally to the earth’s surface via seeps or springs.
group
An artificial category between species and cultivar used to designate a collection of cultivars with similar parentage.
growth regulator
A chemical that sp.s or slows plant growth or maturation.
grub hoe
A heavy weight hoe for digging roots, etc.
guard cells
The two cells that bound a stoma and by opening and closing allow gas exchange.
guard petals
The outer petals of a rose, especially those that are larger than and also enclose the inner petals.
guides
A term applied to the large parenchyma cells seen in the cross-section of the costa of many Dicrana.
gulch
Western U.S. term for a deep cleft, especially one that is formed following a torrent; a ravine. See also: arroyo, gully, valley.
gully (adj. gullied)
A miniature valley eroded by water. See also: ravine, valley.
guttation

The formation of drops of water on plants from moisture in the air. 2. The exudation of liquid water from the uninjured surface of a plant leaf. 3. The process of water being exuded from hydathodes at the enlarged terminations of veins around the margins of the leaves.
gymnosperm
A seed plant that bears naked seeds, i.e., seeds without ovaries. These include conifers, cycads, ginkgos, and ephedras.
gymnospermous
Bearing naked seeds, without an ovary.
gymnostomous
Without a peristome.
gynandrium
A structure in which the stamens are attached to the pistil.
gynobase
An enlargement or prolongation of the receptacle bearing the ovary.
gynobasic
Describes a style arising from the base of the gynoecium.
gynodioecious
Having both bisexual flowers and female flowers, but on separate plants. See also: dioecious.
gynoecium
The pistil or pistils considered as a group.
gynomonoecious
Refers to a plant that produces both perfect flowers and female flowers.
gynophore
A stalk bearing the gynoecium above the level of insertion of the other floral parts.
gynosporangium
The receptacle in which gynospores are developed.
gynospore
One of the larger (female) reproductive bodies in the Isoetaceae, etc.
gynostegial corona
The collective term for the staminal and interstaminal coronas (in Asclepiadaceae), both of which are associated with the gynostegium. See also: staminal corona, interstaminal corona.
gynostegium
The crown of united stamens in milkweeds, Asclepidaceae.
gynostemium
The compound structure resulting from the union of the stamens and pistil in the Orchidaceae.
gypsum
A mineral consisting of hydrous calcium sulfate that is used as a soil amendment to add sulfur and calcium and counteract salt damage. It is added to cement to regulate setting.
h & s
The abbreviation for the height and spread of a plant.
habit
The general appearance of a plant.
habitat
The natural dwelling place of an animal or plant; the type of environment where a particular species is likely to be found.
haft
Narrow stalk-like base to some petals, as those on irises.
hair

An epidermal outgrowth composed of a single elongated cell. 2. In lichens, a multicellular outgrowth from the cortex.
hairy
Covered with hairs; pubescent, hirsute, etc.
half-inferior
Describes an ovary partly below and partly above the level of attachment of the perianth and stamens.
halophyte

A plant adapted to living in highly saline habitats. 2. A plant that accumulates high concentrations of salt in its tissues.
hamate (syn. hamulose)
Curved like a hook.
hand axe
A similar tool intermediate in size between a hatchet and an axe.
hapaxanthic
Reproducing once only at the end of the plant’s life.
haptotropism (adj. haptotropic)
An orientation response of an organism to stimulation by touch.
hard construction (alt. hard landscaping)
The nonliving elements of a landscape installation, made of wood, brick, concrete, etc.
hard frost
A frost where both the air and the soil has dropped below freezing. Many plants can survive a light frostbut cannot survive a hard frost.
harden off
To gradually accustom a plant to more difficult living conditions, e.g., moving a plant from the greenhouse to the partial shade of a tree before planting it in a garden.
hardening

The process of gradually taking plants into a harsher environment, e.g., from the hothouse to the garden. 2. The term can also mean sustaining a plant from summer to winter, which may include a three-staged process: 1) phytochrome clocks signal the shortening days with a color change. 2) Growth ceases, carbohydrates are transported to the roots, and abscisic acid forms at the union of leaf and stem, dropping the leaf and healing the wound. The dropped leaves serve as mulch and protect the roots from excess cold in the winter, while cell walls toughen. 3) A smooth ice forms around the cells without rupturing them, a process called vitrification.
hardiness
The ability of a plant to withstand winter cold and summer heat.
hardpan
A layer of soil sufficiently clogged with clay or other particles which often prevents the penetration of water and shrub or tree roots.
hardscaping
Man-made objects of a landscape as opposed to natural objects like plants and streams.
hardware cloth
A metal fabric that is more coarse than a window screen but finer than fencing.
hardwood
A term applied to broad-leaved trees as opposed to conifers.
hardwood cutting
Cutting taken from a mature woody stem for the purpose of propagation.
hardy
A term used regarding plants that describes their ability to withstand cold. It does not mean that the plant is long-living, pest resistant, or drought tolerant.
harrow
A field implement with semicircular teeth that breaks up clods and levels soil. 2. To break up and level the soil surface by dragging over it an implement of the same name designed for that purpose.
hastate (syn. halberd-shaped)
Like an arrow-head, but with the basal lobes pointing outward nearly at right angles.
hatchet
A short axe.
haustorium
A specialized absorbing structure of a parasitic plant through which it obtains chemical substances from its host.
head

A dense cluster of sessile or nearly sessile flowers on a very short axis or receptacle; heart-shaped. 2. Ovate with two rounded lobes and a sinus at the base; commonly used to define such a base. 3. The number of animals, e.g., 40 horses, that would be counted by a farmer/rancher as 40 head of horses, or merely 40 head.
head back
To cut back the main branches of a woody plant severely.
heading back
Cutting a branch back to a bud or side branch to increase the number of shoots, making the plant thicker and bushier.
heartwood
The harder and often darker colored wood that forms the interior of a tree trunk or branch.
heath (alt. heathland)
An extensive area of rather open uncultivated land usually with poor coarse soil and covered with low shrubs, such as those of the genus Erica; a moor.
heavy soil
An imprecise term which refers to soil in which the particles are packed closely together with little air or water available to the roots of the plant.
hedge
A fence or boundary formed by a dense row of shrubs or low trees and often given a formal appearance by frequent trimming.
hedge shears
Specialized heavyweight scissors for shaping woody plants into hedges.
hedge trimmer
A hedge trimmer is a gardening tool or machine used for trimming (cutting, pruning) hedges or solitary shrubs (bushes). Different designs as well as manual and powered versions of hedge trimmers exist.
hedgerow
A row of shrubs or trees enclosing or separating fields, often on a low berm of earth.
heel
The portion of old wood at the base of a cutting.
heel in
To temporarily store plants with their roots in moist soil or sawdust to hold them for several days or weeks until they are able to be planted properly.
heirloom garden (syn. antique garden)
A garden created using plants that were in cultivation in the past, usually from a particular period.
heirloom plant
A plant that was developed and in cultivation sometime in the past.
helicoid
Having the shape of a flattened coil, often describing a cyme.
heliophilous
Adapted to, or capable of, growing in full sunlight.
hemiepiphytic
Remaining rooted in the ground but climbing tree trunks.
hemiparasite (syn. semiparasite)
A parasite which lives on and derives part of its nourishment from its host, but also conducts photosynthesis; for example, the mistletoe.
hemostatic
Herbal medicine that can stop bleeding.
hemp

The course fibers used to make cordage derived from plants of the genus Cannibis, or other similar plants. 2. A plant of the genus Cannibis.
herb garden
A garden consisting of culinary and/or medicinal herbs and often having an ornamental design.
herbaceous

Having little or no woody tissue; leaf-like in color and texture. 2. Refers to a plant which dies back to the roots each year during winter, as opposed to a plant which remains green all winter.
herbal

A book about herbs, usually illustrated. 2. Of, or relating to, herbs.
herbalism
The cultivation, collection, study and use of herbs, particularly for medicinal purposes.
herbalist
One who practices herbalism.
herbarium

An organized and cataloged collection of plant specimens. 2. A specialized room or building with constant levels of temperature, moisture and restricted light where plant samples are stored in a designated pattern in large light proof cabinets, allowing samples to remain useful and to be retrieved for study and comparison for centuries.
herbarium glue (alt. herbarium paste)
An adhesive which minimizes cracking, discoloration, and shattering with age, used in fastening plant specimens to the herbarium sheet.
herbarium sheet
Lightweight white card stock used as backing for herbarium specimens, in a standard size of 11 X 16 inches (28 X 40 cm.).
herbicide
A substance that is fatal to plants, or to selected plants.
hermitage
A usually small building provided for contemplation, especially of nature.
hesperidium
A relatively large fruit, pulpy inside with a hard rind outside, such as citrus fruits; technically a berry.
heteroblastic
Describes a plant which has adult parts distinctly different in form from the juvenile parts.
heterocarpous
Producing more than one kind of fruit.
heteroecious
Parasitic on alternating hosts; starting life on one organism, then affecting a second species.
heterogamete
Either of a pair of gametes that differ from each other in shape, size, or behavior, usually occurring as large nonmotile oogametes and small motile sperms.
heterogamous
Bearing two kinds of flowers.
heterogamy
The union of recognizably male and female gametes.
heteromallous
Turned in different directions.
heteromorphic (alt. heteromorphous)
With different forms during the life-cycle; e.g., a fern with the sporophyte different in form from the gametophyte.
heterophyllous
The presence of two or more distinct leaf shapes on a single individual.
heterostyly (adj. heterostylous, adj. heterostyled)
A species in which flowers are similar except that the stigmas and anthers are held at different levels relative to each other, because style length differs between plants. See also: homostylous.
heterotrophic
An organism that requires chemical energy from already formed organic molecules; dependent on organic food made by photosynthetic plants.
hexaploid
Having six sets of chromosomes.
high analysis fertilizer
A powder containing large quantities of nutrient elements.
high-centered
Having the central petals longest; the classic hybrid tea rose form.
higher taxa
Those names ranking above the species level, e.g., genus, family, order, etc.
hillock
A small hill. See also: knoll.
hilum
The scar or point of attachment of the seed.
hip
The closed and ripened receptacle of a rose which contains the seed.
hippocrepiform
Horseshoe-shaped.
hirsute
Pubescent with rather coarse or stiff hairs.
hirsutulous
Slightly hirsute.
hirtellous
Minutely hirsute.
hispid
Beset with rigid or bristly hairs or with bristles.
hispidulous
Minutely hispid.
hoary
Grayish-white with a fine close pubescence.
holocoenotic
The theory that environmental factors act as a whole or aggregate in their effect upon organisms.
holotype
One pressed herbarium specimen designated by the author as the plant on which the description and name are based. See also: isotype, lectotype, syntype, topotype, nomenclatural type, neotype.
homeopathic
Herbal medicines using the system of homeopathy founded in the 1700s. The theory is that ‘like cureslike’ so minute doses can cure; for example, acid will cure an ulcer.
homochlamydeous
Refers to a flower in which sepals and petals are so similar that all are called tepals.
homogamous
Able to bear just one kind of flower.
homologous
Refers to organs or parts that are similar in form or function.
homology (pl. homologies)
The study of structural similarities that indicate actual physical relationships in which species with the same ancestors retain many of the same traits. See also: convergence.
homomallous
Turned in the same direction.
homomorphic
Shaped the same.
homonym
A scientific name given two or more times to plants of the same taxonomic rank but which are quite distinct from each other. See also: basionym, synonym, tautonym, autonym.
homosporous
Having spores of a single appearance and behavior.
homostyly (adj. homostylous, adj. homostyled)
A species in which the flowers have stigmas and anthers held at the same level relative to each other on all plants. See also: heterostylous.
honeydew drip
The sticky mess on your car after parking under trees which have aphids or other sucking insects infesting the tree. Certain ants and fungi also feed on the honeydew.
hormone
A biochemical product of a specific cell or tissue that causes a change or activity in a cell or tissue located elsewhere in an organism.
horny
With a texture which is hard and brittle, but having a fine texture and is easy to cut.
hort.
A term used in a botanical name to indicate that it is not a properly accepted binomial, but is a name used by gardeners and nurseries, perhaps published, but in nursery catalogs rather than scientific journals. It can stand for three terms: hort., meaning “of the garden;” hortorum, meaning “of gardens;” or hortulanorum, meaning “of gardeners.” All three are abbreviated hort., and are never capitalized so as to avoid being mistaken for an author’s name.
hortus conclusus
An enclosed garden, as that of a monastery. in art, a representation of the Virgin and Child in a fenced garden; from the latin, “enclosed garden”. See also: hortus deliciarum.
hortus deliciarum
A walled garden, like the hortus conclusus, but geared more toward the pleasure of its ownsers and more secular in its motifs; from the latin, “pleasure garden”.
hortus fenestralis
A window garden; a window box in the form of a miniature greenhouse; from the latin, “window garden”.
hortus siccus
A collection of dried and pressed plants, often arranged in book form; an herbarium; from the latin, “dired garden”.
hose-in-hose
With one perfect corolla inside another.
hot caps
Paper or plastic tent covers used to protect small plants from damaging weather conditions.
hotbed
An outdoor pit enclosed with a glass roof and heated by fermenting material, hot water pipes, or electric cables.
houseplant
A plant which can be grown indoors, usually for decorative reasons.
ht.
Abbreviation for plant height.
humate
A salt or ester of a humic acid.
humectant
A substance which promotes retention of moisture; a moistening agent such as crystals of polymer hydrogel, which take up water, releasing it slowly for the roots of thirsty plants.
humidistat
An instrument for measuring, sometimes controlling, the amount of water vapor in the air.
humidity (adj. humid)
The moisture in air.
humidor
A tightly closed container which holds a constant level of moisture in the interior, such as for long-term preservation of seeds or storage of fruits and vegetables.
hummock
A mound rising above the surrounding land, usually overgrown with vegetation.
humus (adj. humic)
Partly or wholly decomposed vegetable matter.
husk
The outer layer of certain fruits like walnuts Juglans, as an outgrowth of the perianth or involucre.
hyalescent
Translucent.
hyaline
Transparent or translucent.
hydathode
An epidermal structure specialized for the secretion or exudation of water.
hydration
The reaction of cement with water to form a chemical compound.
hydraulic seeding
A method of planting grass seed by spraying it in a stream of water, which may contain other materials such as mulch or plant food.
hydric
Of, or adapted to, an extremely moist habitat.
hydromulch
The process of mixing grass seed with water and mulch for spraying onto bare soil directly.
hydrophyte (adj. hydrophytic)
A plant adapted to growing in water, waterlogged soil or on a substrate that becomes inundated on a regular basis.
hydroponics
Raising plants in a totally soilless environment. Plants are grown directly in water with nutrients added as necessary, usually in a greenhouse or under artificial lights.
hydrotropism
The growth of an organism or a part, such as a root, in response to the presence of water.
hygroscopic

Altering form or position through changes of moisture. 2. Readily absorbing water and thereby altered in form or direction. Hygrometric is sometimes used with a similar meaning. 3. Pertaining to water that is electrostatically bound to the surface of dirt particles and is therefore unavailable to plants.
hylea
The primeval forest.
hypanthium
A cup-like base of a flower to which the stamens, sepals and petals are attached.
hypertensive
Refers to an herbal medicine that can elevate blood pressure.
hypha (pl. hyphae)
The microscopic, multicellular, nonphotosynthetic filaments of fungi and seaweeds.
hypochil
The (often fleshy or otherwise modified) basal portion of the labellum or lip in Orchidaceae.
hypocotyl
The part of the stem of an embryo or young seedling below the cotyledons.
hypocrateriform
Salverform; a tubal flower flaring out into a flat top.
hypodermis
A layer of cells immediately internal to the epidermis.
hypogeal
Of or relating to the emergence of cotyledons below the surface of the ground. See also: epigeal.
hypoglycemic

Refers to herbal medicines that can lower blood sugar. 2. A patient with low blood sugar.
hypogynous
Situated on the receptacle beneath the ovary and free from it and from the calyx; having the petals and stamens so situated.
hypolimnion
The lowest layers of a body of water, below the thermocline, where water does not circulate freely.
hyponasty (adj. hyponastic)
In plant physiology, the state in which more vigorous growth occurs in the lower surface of an organ, such as a young fern frond, causing an upward curvature. See also: epinasty.
hypophysis
A swelling of the seta immediately under the capsule.
hypotensive
Refers to herbal medicines that can reduce blood pressure.
hysteranthous
Refers to plants that have flowers develop before the leaves.
idioblasts
Specialized epidermal cells which produce slime or gum.
illegitimate
Refers to a name published validly, but not satisfying one or more articles of the Code of botanicalnomenclature or the Code of Nomenclature for cultivated Plants.
imbricate
Overlapping, either vertically or spirally, where the lower piece covers the base of the next higher, or laterally, as in the aestivation of a calyx or corolla, where at least one piece must be wholly external and one internal.
imbricated
Closely overlapping each other like the tiles of a roof.
immersed

Covered up; when related to mosses, refers to the capsule when the perichaetial leaves project beyond it. 2. Growing wholly underwater.
immigrant
One who moves or immigrates into an area. An animal is an emigrant when leaving one area, but the same animal is an immigrant when moving into the new area.
imparipinnate
Having an uneven number of pinnae; lacking a terminal pinna. See also: paripinnate.
imperforate
Lacking holes.
impressed
Bent inward, hollowed, or furrowed as if by pressure.
in vitro
Outside the living body and in an artificial environment, as a tissue culture.
incised
Cut sharply and irregularly, more or less deeply.
included
Not at all protruded from the surrounding envelope.
incompatible
Refers to plants that will not form a lasting union at a graft.
incomplete flower
Lacking one or more whorls, i.e., sepals, petals, stamens, or carpels, of the complete flower.
incrassate
Thickened, often relating to cell walls.
incumbent
Describes cotyledons lying with the back of one against the radicle.
indefinite
Inconstant in number or very numerous.
indigenous
Native and original to the region.
indumentum
A massing of fine hairs, glands, or prickles.
induplicate
Having edges or margins folded inward.
indusiate
Bearing indusia.
indusium (pl. indusia)
The thin, scale-like covering of immature sori.
industate
Provided with an indusium.
inerm (alt. inermous)
Unarmed, no prickles, thorns, teeth, etc.
inferior ovary
With the flower parts growing from above the ovary; one that is adnate to the calyx.
inflated
Bladdery; applied to the alar cells of leaves when enlarged much beyond the size of the neighboring cells.
inflexed
Bent inwards.
inflorescence
The flowering part of a plant, and especially the mode of its arrangement.
influent

An organism which sways interactions within a community but is not a dominant species; one which influences. 2. The flow of water from a stream into a body of water, especially subterranean storage. See also: effluent.
infra-
As a prefix, denotes below.
infrageneric
Refers to any taxon of a rank below that of genus, such as species, variety, cultivar, etc.
inframedial
Below the middle.
infraspecific
A taxonomic group within a species, such as a subspecies.
infructescence
The grouping or arrangement of fruits borne on a plant.
infundibuliform (syn. infundibular)
Shaped like a funnel.
inoculant (syn. inoculum)
The material used in an inoculation.
inorganic

Being or composed of matter other than that from a plant or animal. 2. Not arising from natural growth; artificial.
inorganic fertilizer
A powdered or liquid chemical mix of nutrients which does not contain carbon.
insect vectors
Bugs that carry and distribute disease-causing microorganisms.
insectivorous (n. insectivore)
Feeding on insects.
inserted
Attached to or growing out of.
integrated pest management (abr. IPM)
A philosophy of pest management based on the idea of using the least dangerous course first; stresses the use of natural controls, such as insect predators, over the use of chemical pesticides.
integument
An outer covering or coat.
inter-
As a prefix, denotes between or among.
intercalary
Of a meristem situated between the apex and the base.
intercalary growth
Growth in the tissue between the apex and the base.
intercalary inflorescence
An inflorescence that either arises in an internodal position, or one originally terminal but ceasing to be so when vegetative growth subsequently resumes from the stem apex.
interfoliaceous
Between the leaves of a pair, as the stipules of many Rubiaceae.
intermediate

Refers to a grafting scion which is attached at both ends either simultaneously or successively. 2. Refers to hybrids apparently combining the characters of both of their parents in equal measure and standing midway between them.
internode
The portion of a stem between two nodes.
interpetiolar
Describes a stipule located between the petioles of two opposite leaves. See also: intrapetiolar.
interrupted
Describes an inflorescence with the flowers unevenly distributed along the axis, with conspicuous gaps. Also describes the fertile frond of some ferns with the clusters of sporangia similarly arranged.
interspecific
Refers to hybrids between two separate species of the same genus.
interstaminal corona
The fleshy lobes, often connate into a tube, attached to the base of the staminal column in the interstaminal areas of members of the family Asclepiadaceae. See also: gynostegial corona.
intertidal
The shore zone between high and low tides.
intra-
As a prefix, denotes within or inside.
intramarginal
Within and near the margin.
intrapetiolar
Describes a stipule located between a petiole and the stem. See also: interpetiolar.
intricate
Tangled.
introduced
Brought intentionally from another region for purposes of cultivation.
introgression
The formation of a range of intermediate plants by hybridization among parents and progeny.
introrse
Turned inward or toward the axis.
invaginated
Inside a sheath.
invalid
Refers to scientific names published with incomplete information or in an invalid publication.
invasive
Spreading aggressively from the original site of planting.
involucel
A secondary involucre, as that of an umbellet in Umbelliferae.
involucellate
Having an involucre.
involucral
Belonging to an involucre.
involucrate
Having an involucre.
involucre
A circle or collection of bracts surrounding a flower cluster or head, or a single flower.
involute

Rolled inward. 2. With margins of ray florets rolled forward along their longitudinal axis. When fully involute, the margins touch or overlap so only the reverse of the floret is visible.
iron chelate (alt. chelated iron)
A powdered compound used as a soil additive to boost the amount of iron available to plants.
irregular

Varying in form. 2. Asymmetrical.
irregular flower
Describes a flower showing inequality in the size, form, or union of its similar parts.
irrigation
The provision of water to plants to supplement rainfall. See also: drip irrigation, center pivot irrigation.
isidia
Finger-like outgrowths from the upper cortex of a lichen.
isodiametric
Having equal diameters.
isogamy
The union of equal gamete.
isolateral
Having upper and lower surfaces which are similar in form.
isomorphic
Having a similar form, but being genetically different.
iteroparity
Refers to an organism that has multiple reproductive seasons over its lifetime. See also: semelparity.
jointed

With nodes on the stem. 2. Having obvious, thickened areas between cells.
jugate
Paired, with prefixes for the number of pairs; bijugate would be two pairs.
jugum
A pair.
julaceous
Smooth, slender, and cylindric; like a catkin or a worm.
junceous
Rush-like.
juvenile
Young; not adult.
karyoevolution
Changes in the chromosome set of a number or structure occurring through evolutionary change.
karyotype
Characterization of the chromosome set of an individual or group, described in terms of number, length, centromere position, etc.
katabolism
The process of breaking down protoplasm with the liberation of energy and the formation of simplesubstances.
kelp
Any of various large brown seaweeds, sometimes used to enrich poor soil.
kernel
The nucleus of an ovule or seed; everything inside the coat.
kingdom
One of the primary taxonomic categories into which natural objects are commonly classified. There are currently five kindoms used: Plantae, Animalia, fungi, Monera (bacteria) and Protista (protozoans, some types of algae, etc.).
kitchen garden
A garden where vegetables, fruits, and herbs are grown for use in cooking.
knees
When growing in a swamp, the above-water spongy roots of cypress, Taxodium. These provide air to the interior systems of the tree.
knot garden
An elaborately designed garden consisting of flowers, herbs, and/or low shrubs arranged in intricate, geometric, knot-like patterns when seen from above.
knotted
Having nodes or lumps.
krummholz (syn. elfin forest)
Scrubby, dwarfish growth of trees, often forming a distinctive zone at the tree line of mountains.
kwongan
The sclerophyllous vegetation of the sand plains of south-western Western Australia.
labiate
Lipped; belonging to the Labiatae.
labriform
Lip-shaped.
labyrinthiform
With complicated sinuous lines or winding passages.
lacecap
A flattop inflorescence in which the outer flowers are larger than the inner ones, and the outer are sterile.
lacerate
Irregularly cleft as if torn.
lachrymiform (alt. lacrymiform)
Tear-shaped.
laciniate
Slashed; divided into narrow pointed lobes.
laciniated
Describing dahlias with the split or fimbriation in proportion to ray floret length, and no less than 1/6 of ray floret length, there should be a twisting in the area of the split involute or revolute ray florets, to give an overall fringed effect.
lactifer (alt. lacticifer)
A latex duct which may or may not produce the milky sap.
lactiferous (alt. lacticiferous)
Describes a plant with a milky sap; producing latex.
lacuna
A cavity or gap, usually referring to one found in tissue.
lacunose
Pitted.
lacustrine
Refers to, produced by, or formed in a lake. Lacustrine wetlands are those reaching into a lake and acted on by waves and currents.
laevigate
Smooth, polished.
lageniform
Shaped like a bottle or flask.
lagoon
A relatively shallow pond, sound or lake, especially one connected to the sea.
lake plain
The nearly flat bed of an extinct lake or the lowland surrounding an existing lake.
lake senescence
The aging process of a wetland or lake that results from excess sediment and vegetation.
lamella (pl. lamellae)

A thin flat plate or laterally flattened ridge. 2. The acid crystals of lichens shaped in this way.
lamellate (alt. lamellose)
Composed of or arranged in layers or thin plates; having lamellae.
lamina
A blade; the leafy portion of a frond.
laminal
Superficial on the surface of thalli or lobes of lichens.
lanate (alt. lanose)
Woolly.
lance-shaped
Elongate in shape, broadest below the middle and gradually rowed toward the tip.
lanceolate
Shaped like a lance-head, several times longer than wide, broadest above the base and narrowed to the apex.
landlocked property
A parcel of real estate which does not border any road for access.
landraces
Crop cultivars or animal breeds that evolved with, and has been genetically improved by, traditional agriculturalists, but has not been introduced in modern breeding practices.
landscape architect
A person trained in the location of landscape elements for human use and enjoyment. It involves the placement of structures, traffic flow, and plantings.
landscape architecture
The design of outdoor space for maximum enjoyment.
landscape contractor
A company or individual performing landscape installations.
landscaper
An improper term applied to anyone doing landscape work. Often the person is untrained.
language of flowers (alt. symbolism of flowers)
A complex code under which each species of flower, even individual colors of that species, carries a meaning or message.
lanose (alt. lanate)
Woolly.
lanuginose
Nearly lanate, but with shorter hair.
lasion
A periphyton in which the organisms are densely populated and are interdependent. See also: epiphyton.
lateral
Side shoot, bud, etc.
lateral cane
Any cane growing from a main cane.
lath
In gardening, any overhead plant protection structure that reduces direct sunlight or protects plants from frost. Wooden strips called laths, wired together in snow fences, were originally used for that purpose.
latiseptate
Having broad partitions. See also: angustiseptate.
latrorse
Turned sideways.
lawn
An area of cultivated and mown grass.
lawn mower
A lawn mower is a machine utilizing one or more revolving blades to cut a grass surface to an even height. Most common types are the reel mower, walk-behind mower, ride-on tractor and zero turn mower.
leader
The highest terminal shoot of a plant.
leaf
A usually flat, green structure of a plant where photosynthesis and transpiration take place and attached to a stem or branch.
leaf axil
The angle between a petiole and the stem.
leaf blight
Any of various diseases which lead to the browning and dropping of a plant’s leaves.
leaf bud
A bud which contains undeveloped leaves.
leaf curl
A disease that causes leaves to roll up.
leaf gap
A break in a stem’s vascular tissue above the leaf trace.
leaf litter
The leaves that have fallen from a plant, either through normal seasonal changes or due to disease. Especially in the latter case, leaf litter can harbor pathogens and should be cleaned up promptly, particularly around plants such as roses. In a naturalized, woodland setting, leaf litter can be a normal part of the workings of the garden.
leaf miners
Tiny grubs that tunnel in leaves leaving whitish blotches or trails.
leaf mold
A form of humus composed of decayed leaves, often used to enrich soil.
leaf primordium
A lateral outgrowth from the apical meristem that develops into a leaf.
leaf scar
The mark left on the twig at the point of attachment of a leafstalk when the leaf falls.
leaf spot
Any of various plant diseases that cause well-defined areas of tissue to die creating noticeable spots.
leaf trace
The strand of vascular tissue between the vascular bundle of the stem and the leaf.
leaflet
A single division of a compound leaf.
leathery
Of a smoothly tough texture.
lectotype
A specimen selected from the original material used by the author in naming a taxon, when no holotypewas designated or if the holotype is missing. See also: holotype, isotype, nomenclatural type, syntype, topotype, neotype.
lee shore (alt. leeshore)
The shore toward which the prevailing winds are blowing. It is not protected from strong wave action and may have breakers. See also: windward shore.
leeward
The side opposite that from which the wind blows, the sheltered side. See also: windward.
leggy
Refers to a plant growing tall and spindly, usually from inadequate sunlight.
legume
The dry, single-celled fruit of the Leguminosae, formed of a simple pistil, and usually dehiscent by both sutures and often grouped in a long pod, as found in a pea-pod.
leguminous
Pertains to a legume or to the Leguminosae.
lenticular
Lentil-shaped; of the shape of a double-convex lens.
lentiginous
With very tiny dots, as though covered with dust.
lepidote
Beset with small scurfy scales. See also: leprous.
leprous
Scurfy, covered with tiny scales. See also: lepidote.
leptocaul
With a thin primary stem.
leptodermous
Thin-coated; applied to capsules when soft and pliable.
leptosporangiate
Describes a fern with sporangia having walls only one cell thick. See also: eusporangiate.
leptosporangium
A thin-walled sporangium originating from a single epidermal cell.
leucoplast
A colorless plastid in the cytoplasm of interior plant tissues that is potentially capable of developing into a chloroplast.
liana (alt. liane)
Any of various high-climbing woody plants, usually found in the tropics.
lianoid
Vine-like.
lichen
An organism formed by the symbiotic association of an alga and a fungus.
life form
Characteristic structure of a plant or animal.
lifting
Digging up and removing a plant from its surrounding soil.
light frost
A frost where the air has dropped below freezing but the ground has not. Many plants can survive a light frost but cannot survive a hard frost.
light shade

Incomplete shade, where some sunlight is still available. 2. An area in shade for only part of the day.
lignin
The chief noncarbohydrate constituent of wood, a polymer that functions as a natural binder and support for the cellulose fibers of woody plants.
lignotuber
A woody swelling on the trunk of a tree or shrub, below or just above the ground, that contains adventitious buds from which new shoots develop if the top of the plant is cut or burnt.
lignum vitae
The very hard, heavy wood of any of several trees of the genus Guaiacum.
ligulate
Furnished with a ligule.
ligule
A strap-shaped corolla, as found in the rayflowers of Compositae. 2. A thin often scarious projection from the summit of the sheath in grasses.
limb

A branch of a tree. 2. The expanded portion of any petal, or of a leaf, such as the expanded portion of a gamopetalous corolla above the throat. 3. The upper part of a leaf as distinct from the leaf base.
limbate
Describes a leaf bordered by a part of another color.
limbed
Refers to timber with branches removed.
lime-dots
White salt concretions around a hydathode.
lime-hater
Plant unable to thrive in alkaline.
limnology
The study of fresh water bodies with regard to physical, chemical, geological, meteorological, biological, and ecological conditions. Oceanography encompasses the same specialties for marine salt waters.
line-out
To set out young rooted plants in the outdoor nursery to grow larger or on which to work grafts.
linguiform
Tongue shaped.
Linnaean (alt. Linnean)
Of, or relating to, the Swedish botanist Carolus Linnaeus (1707-1778), or the taxonomic classificationsystem of binomial nomenclature he originated.
lip
Each of the upper and lower divisions of a bilabiate corolla or calyx; the peculiar upper (but by a twist of the pedicel apparently lower) petal in orchids.
lip petal
The lower petals of some irregular flowers, often elaborately showy, as in orchids.
lister
An agricultural implement consisting of a double plow, in which the plowshares force the soil in opposite directions, forming a series of alternate ridges and furrows. The basin lister comes with an attachment that forms low soil dams across each furrow at intervals of 15 to 25 feet (to 6.5 m.), forming basins to hold large amounts of water.
literature retrieval
The process of selecting–manually or automated–appropriate documents within the mass of published literature that are of interest to the user. See also: document retrieval, data retrieval.
lithic context (syn. stratigraphic context)
The rock layers surrounding a fossil that allow the age of the fossil to be determined.
lithophyte
A plant that grows on rocks.
littoral
Of or existing on a shore.
littoral drift
Sediment and detritus moved along the shore by waves and currents. See also: littoral zone.
littoral zone
The shallow water along the shore from zero to the depth where plants no longer root, about 10 feet (3 m.) deep.
liverwort
A nonflowering plant in the botanical class Hepaticae, closely related to the mosses.
loam (syn. loamy soil)

A generally fertile and well-drained soil, containing clay, sand, and a significant amount of decomposed organic matter. 2. Any soil.
lobate
Having numerous lobes.
lobe
Any segment of an organ, especially if rounded.
lobedleaf
A leaf whose margin is shallowly divided.
lobulate
With irregular, rounded lobes.
location survey
A less expensive land survey done to only prove that a specific building or right of way is situated on a plot of land, rather than the more detailed metes and bounds survey which gives all border limitations and positions.
lociation
A variation of a climax association, differing from the normal in the kinds of subdominants. See also: faciation.
loculicidal
Longitudinally dehiscent between the partitions of the locule, as in Liliaceae. See also: septicidal.
locus (pl. loci)
The position in a chromosome of a particular gene or allele.
lodicule
One of a pair of tiny scales at the base of a grass floret, believed to a be reduced perianth segments.
loess
A deposit of relatively consistent fine soil, mostly silt, usually carried by wind.
log lifter
A tool designed to easily roll a log, a cant hook, or pike.
loment
On legumes, a pod which narrows between the seeds, drying and splitting apart at maturity into one-seeded segments.
lomentum
A pod constricted between the seeds.
long-day
Of, relating to, or being a plant which flowers only after being exposed to daylight for periods longer than a particular length, usually in the late spring or early summer.
long-lived
Describes a plant where the individuals subsist for a number of years.
longitudinal
Lengthwise.
loppers (alt. lopping shears)
Long-handled pruning shears for cutting larger branches; may be rachet tools, with telescoping handles or specialized long reach type.
lorate
Strap-shaped.
lotic
Of, relating to, or living in actively moving water. See also: lentic.
luculicidal
Dehiscent into the cavity at a cell through the dorsal suture.
lumen (pl. lumina)
The central cavity of an organ or cell.
lunate (alt. lunulate)
Of the shape of a half-moon or crescent.
lustrotts
Glossy.
lutein (syn. xanthophyll)
A yellow carotenoid pigment, usually found with chlorophyll in plants. Also called xanthophyll.
lyrate
Pinnatifid with a large and rounded terminal lobe and with the lower lobes small.
machete
A large powerful knife used to clear brush, e.g., trails through jungle areas.
macrophytes
All visible plants in any habitat, not microscopic bacteria or algae.
macrosporangium
The receptacle in which macrospores are developed.
macrospore
The larger kind of spore in Selaginellaceae, etc.
maculate
Blotched with broad, irregular patches of color.
main shoot
A basal cane or a strong lateral cane.
male cone
The conical, pollen-bearing male element of a conifer.
mallee
A growth form in which several stems arise from a lignotuber.
malpighiaceous hairs
Hairs which are straight and oppressed but attached by the middle.
mamillate (syn. mammillar)
Convex with a short projection in the center.
mammillate (alt. mammillated)
Shaped like a nipple.
manila coin envelopes
Small easily labeled containers for otherwise easily lost parts of plant specimens such as seeds, individual flowers, leaves, etc.
manure
Animal dung used as a fertilizer.
marcescent
Withering but persistent.
marginal rays
Fully developed ray florets which establish and determine the maximum diameter of a flower, as distinguished from the immature central rays (used in describing dahlias.)
marginate
With a border of a different color.
maritime
Describes climate mainly influenced by the sea, generally with mild winters, cool summers, and frequent rain.
marl
A mixture of clay and the carbonates of calcium and magnesium, from precipitation in acidic waters, and from dissolved shells and limestone.
massula (pl. massulae)
Structures associated with megaspores or enclosing and trapping microspores to increase the likelihood of fertilization.
matted
Formed into a mat.
mattock
A waist-high tool with a metal head consisting of a horizontal blade with one end twisted as a dull axeand the other end flat or pointed. It is used for breaking hard soil, digging roots, and other heavy work.
maze (syn. maze garden)
A confusing intricate network of passages, particularly one created with tall, dense hedges forming the walls and separating the pathways.
meadow
Typically, a level grassland or field within a larger ecosystem, such as a forest. Often, the grass grown on its natural, low-lying, moist areas is used for forage or fodder or cut for hay.
mealy
Farinose.
mealybug
Any of the scale insect belonging to the family Pseudococcidae that have a white powdery covering and are destructive plant pests, particularly toward fruit trees.
median
Refers to the central crosswise area of a leaf, as ‘median width’ would be the broadest width at midpoint.
median leaf-cells
Cells from the middle of the leaf.
medicinal herb
Any herb which is used as a curative or preventative.
medifixed
Attached by or at the middle.
medulla

The central tissue of a structure, generally referring to the pith. 2. The inner part of the thallus of a lichen.
medullary
Made up of pith, spongy.
megasporangium (pl. megasporangia)
The female sporangium containing the megaspores.
megasporophyll
The leaf bearing the megasporangia.
meiosis
Cell division; the nuclear division that halves the chromosome number.
meiotanglum
The sporangium or gametangium in which meiosis occurs.
mellitophily
Pollination by bees.
membranaceous (syn. membranous)
Like a membrane; thin, rather soft, and more or less translucent.
membranous (alt. membraneous, alt. membranaceous)
Thin in texture, soft and pliable.
meniscoidal
Thin and concavo-convex, like the crystal of a watch.
mentum
A chin-like extension on a flower, particularly in Orchidaceae.
mericarp
One of the achene-like carpels of Umbelliferae.
meristem cloning (n. mericlone)
Artificial propagation of a plant using cells taken from the meristem of a parent plant and yielding genetically identical offspring.
Merrill cases
Cardboard containers purchased flat for easy transport, then opened in the field when needed to hold plant specimens.
mesic
Of, or adapted to, a temperate, moderately moist habitat; neither xeric nor hydric.
mesocarp
The fleshy, middle portion of the wall of a succulent fruit between the skin and the stony layer.
mesophyll
The middle and photosynthetic tissue of a leaf.
mesophyte
Dryland plants, xerophytes, have adaptations to survive for months with no water; water plants, hydrophytes, live in water; mesophytes are all other plants–which need small amounts of water on a regular basis.
mesosperm
The middle layer of the coat of a seed.
metaxylem
The primary xylem formed after the protoxyle.
metes and bounds
A surveyor’s term meaning the legal circumscribing lines setting the outer limits of a tract of land.
microclimate
Climate specific to a small area; may var. significantly from that of surrounding areas.
microconidia
Bacilliform cells produced in pycnidia.
microenvironment
The environment of a very small area.
microevolution
Evolution as a result of very small genetic variations, which may cause the formation of a new subspecies.
microfossil
A microscopic fossil, e.g., that of a pollen grain.
microgametophyte
The male gametophyte, formed by vegetative growth of the microspore of a heterosporous plant.
microhabitat
A very small habitat, such as a rock crevice or a clump of grass.
micronutrient
An organic compound, such as a vitamin, essential in minute amounts to the growth and health of a living organism.
microphyll
A very small leaf.
micropropagation
Propagation of plants through tissue cultures.
micropyle (adj. micropylar)

A minute opening on the ovule through which the pollen tube usually enters. 2. The point upon the seed at which was the orifice of the ovule.
microsporangium (pl. microsporangia)
The male sporangium in which microspores are developed.
microsporophyll
The leaf bearing the microsporangi.
midrib
The central or main rib of a leaf.
midvein
The principal vein in the blade of leaf, pinnule, or segment.
mildew
A fungus that leaves a thin white coating on the surface where it grows.
mimosiform (alt. mimosoid)
With round or conical flower heads having inconspicuous sepals and petals but conspicuous stamens, as found in the genus Mimosa.
ming
A dwarfed evergreen conifer grown as bonsai.
minimum viable population
The isolated population of least numbers which has a good chance of surviving despite the foreseeable effects of demographic, environmental, and genetic events and natural catastrophes.
misapplied
Refers to the usage of a taxon name for an incorrect plant; such names are homonyms and also synonyms of the plants to which they truly describe.
mitochondrion (pl. mitochondria, syn. chondriosome)
Cytoplasmic structure containing enzymes used in converting food to energy.
mitosis
Cell division in which the number of chromosomes in the daughter cells is the same as that of the parent cell. See also: meiosis.
mitriform
Shaped like a mitre or cap.
moat
A ditch filled with water and usually acting as a barrier.
moniliform
Resembling a string of beads; cylindrical with contractions at intervals.
mono-
A prefix meaning one.
monocarpellary
Refers to fruit with one female organ.
monocarpic
Describes a plant that dies after flowering just once.
monocephallic (alt. monocephalous)
Bearing a single flower head.
monocephallous
Having a single flower head.
monochasial cyme
Like a dichasial cyme, but with branches on only one side.
monochasium (pl. monochasia, adj. monochasial)
A cyme with a single flower on each axis of the inflorescence.
monochlamydeous
Describes a flower which has a single whorl of perianth parts.
monoclinous (alt. monoclinus)
Having pistils and stamens in the same flower.
monocolpate
Refers to pollen grains with a single furrow.
monocotyledon (alt. monocot, adj. monocotyledonous)
A plant with one cotyledon or seed leaf.
monoculture
The agricultural practice of planting a field or other land mass with a single crop, all of the same age, like wheat or pine trees. Single crop planting often leads to increased infestation by disease or insects.
monoecious (alt. monoicous)
Having stamens and pistils in separate flowers on the same plant.
monoembryonic
Refers to seeds or ovules with a single germ cell or embryo.
monogeneric
Refers to a family or taxon of higher rank made up of a single genus.
monolete
Describes spores having a single, unbranched scar.
monomial
The single word designation such as genus Iris, or family Iridaceae. See also: binomial.
monopetalous
In the strictest sense, refers to a flower with a single petal; however, it is often used to refer to gamopetalous flowers in which several petals are united to form a tube.
monophyletic
Descended from a single ancestral line. See also: polyphyletic.
monoploid
Having a single set of chromosomes.
monopodium (pl. monopodia, adj. monopodial)
The main axis of a stem or rhizome maintaining a single direction of growth and giving off lateralbranches or stems. See also: sympodium.
monospecific
Refers to a genus which has a single species.
monotypic
A taxonomic division that has only one subdivision, as a family containing only one genus, or a genus with a single species.
montane
Of, growing in, or inhabiting mountain areas.
morphocline
A graded series of character states of a homologous character.
morphology
The study of the form and structure of an organism.
moss
A small nonflowering plant of the class Musci.
mother bulb
A mature bulb which has produced small offset bulbs.
mother plant

A mature plant from which cuttings are taken. 2. The female ancestor of a hybrid.
mother-cell
The cell that gives rise to a particular structure or particular reproductive units.
motile
Actively moving or capable of moving spontaneously.
mottled
With variegated coloring.
mound layering
A method of propagation whereby a branch or stem is scored and then brought into contact with the soil to spur rooting.
moxa
A Chinese herbal process of burning dried herbs on or above the skin to stimulate an acupuncture point or serve as a counterirritant.
mucilage (adj. mucilaginous)
A viscous, slimy material exuded by certain plants.
mucous
Slimy; viscous.
mucro
A short and small abrupt tip.
mucronate
Refers to a leaf or other organ ending suddenly with a stiff spine as a continuation of the midrib; tipped with a mucro.
mucronulate
Diminutive of a mucronate, with a small spine.
mud flats
The bare level bottoms of bodies of water exposed by a drop in the water level. A mud bar may be exposed in a constant water level by a buildup of sediments.
mud logger (mud logging)
A person who works with geologists, lubricating the cutters of cores, and analyzing the resulting rock fragments for stratigraphy, keeping a record or log of the results.
muddled center
A flower center with petals that are disorganized, not forming a pattern. A term applied to Old Garden Roses.
mulch
An organic or inorganic soil covering, used to maintain soil temperature and moisture and to discourage the growth of weeds.
multi-
A prefix meaning many.
multicellular
Formed of two or more cells.
multicipital
Many-headed.
multifid
Cleft into many lobes or segments.
multigeneric
Refers to hybrids with more than two generations in their ancestry, e.g., x Potinara (Brassavola x Cattleya x Laelia x Sophronitis).
multiline
A term used by nurseries to indicate that a horticultural variety derives from several closely related lines.
multiplanar
Describes divided leaves, with the lobes held in several to many planes.
multiple fruit
One in which the carpels of several flowers join in a single fruit, like a fig, Ficus. This differs from an aggregate fruit which derives from the multiple carpels of a single flower, e.g., a raspberry, Rubus.
multiseptate
Bearing more than two septae.
muricate
With a rough surface composed of many short, hard points.
muriculate
Very finely muricate.
muriform
Divided into many chambers.
muscariform
Shaped like a broom.
mutant

An individual produced with sudden and marked differences from the parent, and with a new genetic pattern. 2. A sport.
muticous
Not pointed.
mutualism
A type of symbiosis in which both members depend on each other for their nutrients or other services.
mycelium (pl. mycelia)
The mass of interwoven filamentous hyphae that forms especially the vegetative portion of the thallusof a fungus.
mycology
The study of mushrooms and other fungi.
mycorrhizal
A term describing plants which have a symbiotic relationship with the mycelium of a certain fungus.
myrmecophilous
Strictly, ant-loving; refers to plants which have symbiotic relationships with ants.
myrmecophyte
A plant in symbiosis with ants.
named cultivar
A cultivar that has been given a recognized horticultural name.
napiform root
A taproot which is broader than it is long, like a turnip.
narcotic
A drug used to relieve pain and induce sleep.
native plant
A plant occurring naturally in an area and not introduced by man; indigenous.
natural layering
The spontaneous rooting of stems when they make contact with the soil.
naturalize (alt. naturalise)

To cause a plant to become established and grow undisturbed as if native. 2. The establishment of exotic species in the wild that can reproduce without human intervention.
navicular
Boat-shaped.
nec
In nomenclature, this is placed between the names of two authors, indicating that neither named the taxon in a valid manner.
neck
The lowest part of the capsule just above the point where it joins the seta.
necrosis
The localized death of living tissue usually caused by a pathogen.
nectariferous
Producing nectar.
nectarivore
An animal, including insects and bats, that eats nectar.
nectary
Any place or organ where nectar is secreted.
needle
The very long and narrow leaf of pines and related trees.
needle-like
Very long, narrow, and pointed at the tip.
nematode
Any of several unsegmented, elongated cylindrical worms of the phylum Nematoda. They may be parasitic in animals or plants, or free-living in soil or water. As parasites, they can cause damage to and even kill plants. Others can be benficial.
neotenic
Refers to retaining juvenile characteristics throughout the life span.
neoteny
The process of fulfilling a function in an imperfect or young state.
neotype
A specimen selected to serve as the nomenclatural type when the material used to base the name of the taxon is missing. See also: holotype, lectotype, nomenclatural type, syntype, topotype, isotype.
nervine
An herbal medicine with a soothing and mildly sedative action.
nest-fronds
Specialized, shield-shaped, basal fronds in some ferns which accumulate leaf litter forming a nest-like covering.
net-veined
With a network of veins.
neuter (syn. neutral)
Without stamens or pistils.
neutral soil
Soil having a pH of 7 and therefore neither acidic nor alkaline.
neve
Granular compacted snow at the head of a glacier, or similar snow elsewhere. See also: firn.
niche

Ecological niche–the role of a plant or animal in the environment, its relationship to all the living and nonliving things around it, e.g., a bat is the nighttime hunter of airborne insects. 2. Habitat niche–the specific part or smallest segment of a habitat occupied by an organism, e.g., a pillbug lives under moist rocks or detritus. See also: biotope.
nigrescent
Turning black.
nitrification
The oxidation, as by bacteria, of ammonium to nitrites and the further oxidation of nitrites to nitrates.
nitrogen carrier
A material such as a commercial fertilizer that contains the essential mineral nitrogen.
nitrogen fixation
The conversion of atmospheric nitrogen into an organic form usable by plants and other organisms. This can be through the agency of soil microorganisms, particularly rhizobia living in nodules on the roots of legumes, or by industrial methods.
nocturnal
Active only at night; flowering only at night.
nodose
Knotty or knobby.
nomen (pl. nomina)
Literally, “name”, particularly a scientific name assigned according to the Codes of Nomenclature.
nomen ambigua (abr. nom. ambig., alt. nomina ambigua)
A name that has long referred to different taxa and should be abandoned.
nomen confusum (abr. nom. confus., alt. nomina confusa)
A name based on a type, the herbarium specimens later proving to be two or more taxa.
nomen conservandum (abr. nom. cons., alt. nomina conseranda)
A taxon name that has been formally accepted under the International Code for botanicalnomenclature as the correct name contrary to the usual principles of botanical nomenclature.
nomen illegitimum (abr. nom. illegit.)
A name that was validly published, but contravenes the International Code for botanical nomenclature; A name which was superfluous or had already been applied to another plant.
nomen novum (abr. nom. nov.)
A name proposed to replace a name which has been rejected.
nomen nudum (abr. nom. nud., alt. nomina nuda)
Literally, “a naked name”. Used in nomenclature to refer to the fact that the name was published without a description or diagnosis and is therefore not acceptable under the International Code for botanical Nomenclature.
nomen rejiciendum (abr. nom. rejic., alt. nomina rejicienda)
Literally, “a name to be rejected”. Used in nomenclature when a decision has been made to conserve an older name and the newer one is rejected. See also: nomen conservandum.
nomenclatural synonym
Any one of two or more different names based on the same plant specimen.
nomenclature
A system or set of terms or symbols; the system of providing taxonomic names for organisms.
non
In nomenclature, it means ‘not’ as in “Artemisia lanata Willd. non Lam.”—when Willdenow in 1823 gave a new plant the same name that Lamarmarck had used in 1783 for a different plant.
nongovernmental organization (abr. NGO)
A nonprofit group or association organized outside of institutionalized political structures to gain particular social objectives such as environmental protection.
nonvascular
Refers to organisms with no tissues or vessels to carry water, minerals, etc., such as mosses, fungi, algae, lichens, etc.
nostoc
A blue-green alga with filaments formed of chains of cells.
notch
A v-shaped indentation.
nothomorph
A term used in taxonomy to designate different hybrid forms derived from the same parent species, ranking essentially as a variety.
novirame
A flowering–later fruiting–shoot arising from a primocane.
noxious weeds
Plants which may cause harm to collectors, such as poison ivy, Toxicodendron radicans; or invasiveexotics or parasites and their host plants which may harm the ecosystem or agriculture of an area. See also: quarantine.
nucellus (adj. nucellar)
The central body of the ovule that encloses the female gametophyte; equivalent to the megasporangium.
nucleolus
A special body in the nucleus.
nucleus (pl. nuclei)

The germ cell of the ovule, which by fertilization becomes the seed; the kernel of a seed. 2. The part of the protoplasm that contains the chromosomes.
nullipores
Calcareous red seaweed.
nurse grass
A fast-growing temporary grass included in most grass seed mixtures to give rapid coverage and protect permanent grasses from soil erosion, sunscald, etc.
nurse tree
A fast-growing tree planted with, or left in place near, a slow-growing, permanent tree in order to provide shade and wind protection for the permanent tree until it has reached useful size.
nut
A dry, hard indehiscent 1-celled and 1-seeded fruit, though usually resulting from a compound ovary.
nutant
Nodding, usually referring to the entire inflorescence rather than a single flower; cernuous.
nutlet
One of several small, nut-like parts of a compound fruit; a diminutive nut.
o.c.
On center; refers to the spacing on landscape plans of materials to be planted.
ob-
A prefix meaning inverted, e.g., obcordate describes a heart-shaped leaf attached at the point rather than at the cleft.
obcompressed
Compressed dorso-ventrally instead of laterally.
obconically
Inversely conical, having the attachment at the apex.
obcordate
Inverted and heart-shaped; e.g., a heart-shaped leaf with the pointed end toward the stem.
obcuneate
Like cuneate, but with the point of attachment at the broad end.
obelisk
An upright four-sided usually monolithic pillar that gradually tapers as it rises and terminates in a pyramid, often inscribed with words or designs.
obhastate
Like hastate, but with the triangular lobes at the tip.
oblanceolate
Lanceolate with the broadest part toward the apex.
oblate
Shaped like a sphere which is depressed at the poles.
obligate
Incapable of surviving without a host, as occurs with certain parasites. See also: facultative.
oblique
Unequal-sided or slanting.
obloid
Having an oblong shape but with the cross section circular and the ends rounded.
oblong
Longer than broad and with nearly parallel sides.
obovate
Inverted ovate; egg-shaped, with the broadest part above, or away from the stem.
obovoid
Having the form of an inverted egg.
obpyramidal
Shallowly triangular with the attachment at the point rather than the middle of the flat side.
obpyriform
Inversely pear-shaped, with the attachment at the narrow end; turbinate.
obscure
Hidden.
obsolescent
Becoming rudimentary; refers to an organ no longer functional and reduced to vestigial remains.
obsolete
Not evident; rudimentary.
obtuse
Blunt or rounded at the end.
ocellated
Having spots.
ochrea
The sheath around the base of the seta, terminating the vaginula.
ochroleucous
Yellowish-white.
ocrea (alt. ochrea)
A legging-shaped or tubular stipule.
octoploid
Having eight sets of chromosomes.
odd pinnate
A pinnate leaf with a terminal leaflet, hence having an odd number of leaflets.
odoriferous
Producing an odor, often fragrant.
officinal
Of the shops; a plant used in medicine or the arts.
offset
A short sideshoot arising from the base of a plant; also a small bulb arising from the base of another bulb.
offshore wind
Blowing from the land across the water, usually warmer than an onshore wind, and with lower waves.
Old Garden Rose (abr. OGR, syn. Old English Rose)
A hybrid rose which has been in cultivation since before 1867.
oleaginous
Refers to fleshy, oily tissue.
oligo-
A prefix denoting few, as oligopetalous means with few petals.
once-compound
A compound leaf with a single set of undivided leaflets.
onshore wind
Blowing from the water to the shore, building large waves as it uses its full fetch. This is usually cooler than an offshore wind.
ontogeny
The course of development of a single organism.
oocyte
A cell from which an egg develops.
oogamy
Conjugation between sperms and egg.
oogonium
The organ producing the egg or eggs.
ooshere
The egg cell or ovum found in the base of the archegonium.
oospore
The fertilized nucleus or germ-cell of the archegonium in cryptogams, from which the new plant is directly developed.
opaque
Dull; neither shining nor translucent.
open pollination
Pollination by natural mechanisms such as insects, wind, etc., as opposed to selective pollination by a plant breeder.
operculate
Furnished with a lid.
operculum
A lid; the upper portion of a circumscissile capsule, which on detachment permits the spores to escape.
opposite
Describes leaves arranged along a twig or shoot in pairs, opposite each other at a single point along an axis.
orangery
A many-windowed building used to house potted orange trees during winter.
orbicular
Circular.
orbiculate
Circular in outline.
orchard
A planting of fruit or nut trees.
order (alt. ordo)
A category of taxonomic classification ranking above the family and below the class.
organelle
A membrane-bound body analogous to an organ and found in the cytoplasm of the cell that performs specific cellular functions.
organic
Derived naturally, from living or once-living matter.
ornamental
A plant that is grown for visual display.
ornamental plants
Ornamental plants are plants that are kept, grown and used for display. Popular in gardens, front yards, homes and businesses alike.
ornamental tree
Generally, a small tree as opposed to a tall, or shade, tree. Often they are flowering trees used as understory plantings, or massed in the open for color and texture.
orthodox seed
Seed that can be dried to moisture levels between 4 and 6 percent and stored without spoiling.
orthotropic
Tending to grow or form along a vertical axis.
orthotropous
Describes an ovule or seed that is erect, with the orifice or micropyle at the apex.
osmosis (adj. osmotic)
The diffusion of liquid through a semipermeable membrane (such as a cell wall) until there is an equalconcentration on both sides of the membrane. The process by which water is taken up through the roots of a plant and transported throughout the structure.
osseous
Bony and brittle.
ostiole (adj. ostiolar)
A small opening or pore.
outbreeding
The interbreeding of distantly related or unrelated individuals.
outcross

A cross between relatively unrelated individuals. 2. The offspring of an outcross.
oval
Egg-shaped. See also: ovate, oviform.
ovary
The part of the pistil that contains the ovules.
ovate
Egg-shaped, with the broader end at the base. See also: oval, oviform.
overhead watering
Watering from above, usually with a sprinkler or a hose with a sprinkler attachment.
overstocked

Refers to a stand that contains more trees or other materials than the site can support, causing tall, weak growth and the tendency to blow down. 2. Refers to a range which has more wildlife or domestic stock than the area can support. See also: fully stocked, understocked, carrying capacity.
overtopping
When one branch of a dichotomy grows more than the other.
overwinter

To survive the winter. 2. To keep alive through winter, e.g., bringing a nonhardy plant indoors, wrapping roses or mulching pansies.
ovid
Broadly elliptical.
oviform
Shaped like an egg. See also: oval, ovate.
oviparous
Producing female reproductive cells.
ovoid
A solid with an oval outline.
ovule
The body which, after fertilization, becomes the seed.
ovuliferous
Bearing ovules.
pachycarpous
Describes a seed having a thick outer layer or pericarp, e.g., the skin on a cherry. The skin (epicarp, ectocarp, or exocarp), flesh (mesocarp or sarcocarp), and stony outer layer of the seed (endocarp) forma pachycarpous covering of the seed proper.
pachycaul
With thick or massive primary construction.
pachydermous
Thick-skinned; applied to the walls of capsules or to cells when firm and resisting.
pack ice (alt. ice pack)
A large area of ice driven closely together.
pagoda
A tower of Far Eastern design usually with roofs curving upward at the division of each of several stories and erected as a temple; a similarly styled garden structure.
paired
Refers to flowers or leaflets in opposite pairs, but bi-, tri-, and multijugate further describe leaflets with two, three, or many such pairs or parts.
palate
A rounded projection of the lower lip of a personate corolla, closing the throat.
palea (syn. palet)
The upper bract which, with the lemma, encloses the flower in grasses.
paleaceous
Clothed with chaff.
Paleobotany (syn. archeobotany)
The study of plant fossils.
paleophytological
Relates to the study of fossil plants.
palisade
Refers to the layer of columnar photosynthetic cells in the leaf.
palm
Any of various plants of the family Palmae, mostly tropical or subtropical monocotyledonous trees, shrubs, or vines, usually having a simple stem and a terminal crown of large pinnate or fan-shaped leaves.
palmate (adj. palmately)
Describes a leaf that is radially lobed or divided.
palmate-veined
With the principal veins arising from the end of the leafstalk and radiating toward the edge of the leaf.
palmately compound
Having veins or leaflets arranged like the fingers on a hand.
palmatifid
Describes a leaf which is deeply, but not completely, divided into several lobes.
palmatisect
Intermediate between palmate and palmatifid.
palustrine
Refers to areas that are not part of a major lake, but are filled with partially decomposed plant materials to considerable depth in wetlands such as bogs, swamps, and marshes. See also: peat.
palustrine wetlands
Those not a part of a main lake, therefore not subject to intensive wave or current effects.
palynology
The scientific study of pollen and spores.
pampas
An extensive, generally grass-covered plain of temperate South America, located east of the Andes and mostly in Argentina.
pan
A layer of different soil below the surface, often impervious, which would be called hard pan.
panacea
A remedy for all ills or difficulties; cure-all.
pandurate (syn. panduriform)
Fiddle-shaped; with a rounded base, narrow waist, rounded upper part and long neck.
panicle
A loose, irregularly compound inflorescence with pedicellate flowers.
panicled (alt. paniculate)
Borne in a panicle; resembling a panicle.
pannose
Like felt in texture, densely covered with hairs.
pantoporate
Describes a pollen grain having rounded apertures all over the surface.
pantropical
Spanning tropical regions around the world.
papilionaceous
Describes a corolla having a standard, wings, and keel, as in the peculiar corolla of many Leguminosae.
papilla (pl. papillae)

A minute, nipple-shaped projection. 2. Small bumps appearing on either the upper cortex or lower surface of various lichens.
papillaer
Minute rounded or acute protuberances.
papillate
Bearing papillae.
papillose (syn. scabrose)
Bearing minute nipple-shaped projections; rough with papillae.
pappus
The modified calyx-limb in Compositae, forming a crown of various character at the summit of the achene.
papule
A relatively large pustule, papilla, or pimple.
papyraceous
Paper-like.
paraclade
An inflorescence on a lateral axis which repeats the symmetry of the primary axis.
paradichlorobenzene
A white crystalline compound used in moth balls and moth crystals and often placed with herbarium specimens to prevent destruction by moth larvae. Large quantities in a closed container may be used to kill insects needed as part of sample.
parallel-ribbed (syn. parallel-veined, syn. parallel-nerved, syn. penniparallel)
With the veins running more or less parallel toward the tip of the leaf.
paraphyllia
Minute leaf-like or much-branched organs among the leaves.
paraphysis (pl. paraphyses)

Jointed hyaline hairs growing among the reproductive organs. 2. The thread-like hyphae between the asci.
parasite
A plant deriving its nutrition from another organism; the dependent member of parasitism.
parasitization
The process whereby one organism lives on or with another as a parasite.
parenchyma
Soft tissue of cells with unthickened walls.
parenchymatous
Cells with broad ends abutting on each other, not dovetailing into each other.
parietal
Borne on or pertaining to the wall or inner surface of a capsule.
paripinnate
Having an even number of pinnae. See also: imparipinnate.
parkland
Land in which clumps of trees are scattered throughout a grassland. See also: parks.
parks
Areas set aside by government and kept in their natural states for the purpose of conserving unique areas for preserving game, walking, riding, or recreation.
paroicous
Having its male and female organs in the same cluster, but not mixed, the antheridia being in the axils of the perichaetial bracts below the archegonia.
parterre
A formal garden of intricately designed, geometrically shaped beds of flowers, herbs, and low shrubsseparated by pathways and/or lawns.
parthenocarpic
Refers to the fruiting of plants which have not been pollinated or otherwise fertilized.
parthenogenetic
Developing without fertilization.
partial
Of secondary rank.
partite
Parted.
partitioned
The pith divided crosswise by woody plates, usually near the leaf scars.
pasture
A grassland which is used as graze for domestic animals.
patellate (alt. patelliform)
Round and thick, with one side concave and the other following the same curve by being convex, like a thick contact lens.
patent (alt. patulous)
Spreading, as do the branches of a tree; particularly at an angle of 26-45 degrees.
pathogen
A disease-causing agent, especially a bacterium, fungi, or other microorganism.
pathological
Diseased.
patina
The changed color or texture of a surface due to weathering or aging.
patulous
More widely spreading than patent.
pea-like
Refers to a flower with a top banner or standard, similar to legume blossoms; papilionaceous.
peat bog
An area with a wet, spongy, acidic substrate, consisting mainly of accumulated layers of sphagnum mossand upon which other plants, and even trees, can grow.
peat moss
Partially decomposed sphagnum moss, often added to soil to increase moisture retention.
peavey (alt. peavy)
A tool similar to a cant hook, but with the tip outfitted with an iron spike rather than a ring.
pectin (adj. pectic)
Any of various water-soluble substances that bind adjacent cell walls in plant tissues and yield a gelwhich is the basis of fruit jellies.
pedicel

A tiny stalk; the support of a single flower. 2. The stalk of the sporangia. See also: seta.
pedicelled (alt. pedicillate)
Borne on a pedicel.
peduncle
A primary flower stalk, supporting either a cluster or a solitary flower.
peduncled (alt. pedunculate, alt. peduncular)
Borne upon a peduncle.
pegging
Securing the ends of canes to the ground so that the plant grows horizontally.
pellucid
Clear, transparent.
peloria (adj. peloric)
Unusual regularity or symmetry in the form of a flower that is normally irregular.
peltate

Describes a leaf attached to the petiole from near the center of the lower surface, and not at the margin. 2. Shield-shaped.
pendant
Hanging downward.
pendular
Having the movement of a pendulum.
pendulous
More or less hanging or declined. Pendulous ovule: one that hangs from the side of the cell.
penicillate
Tufted with small hairs.
penniribbed (syn. penninerved, syn. penniveined)
Having conspicuous lateral veins which are divergent from the midrib and approximately parallel to one another.
pentamerous
Describes a flower which has five parts in each floral whorl.
pentangular
With five angles, as the cross-section of a stem.
pentaploid
Having five sets of chromosomes.
pepo
A fruit, such as that of a melon or squash, having a firm rind, fleshy pulp, many flattened seeds, and a single locule.
perched water table
The water table of a smaller body of groundwater body situated above the general groundwater table.
percurrent
Describes a costa that reaches to the apex of the leaf, but not beyond.
perennate (n. perennation)
To live for more than one growing season, but usually with reduced growth or dormancy between growing seasons.
perennial
A plant whose life cycle lasts for three or more seasons; Lasting year after year.
perfect
Describes a flower having both pistil and stamens.
perfoliate
Describes a leaf having the stem apparently passing through it.
perforate
Pierced with holes.
pergola
An open structure usually consisting of parallel columns supporting an open roof of cross rafters on which climbing plants are trained to grow; an arbor.
perianth
The floral envelope, consisting of the calyx and corolla (when present), whatever their form.
pericarp (syn. fruit wall)
The wall of the matured ovary.
perichaetium (pl. perichaetia, adj. perichaetial)
A whorl of bracts at the base of reproductive organs. In mosses, those surrounding the archegonia and base of seta.
pericycle
The tissue of the stele lying just inside the endodermis.
periderm
An outer, cortical protective layer of many roots and stems that typically consists of phellem, phellogen, and phelloderm.
perigone
The perianth, especially when made up of tepals, or in reference to anything surrounding the reproductive structure.
perigonium (pl. perigonia, adj. perigonial, syn. perigone)

The perianth. 2. In mosses, those bracts surrounding the antheridia.
perigynium
The inflated sac which encloses the ovary in Carex.
perigynous
Adnate to the perianth, and therefore around the ovary and not at its base.
periodicity
Repetition of events at fairly regular intervals.
peripheral
On or near the margin.
periphyton
The assemblage of organisms submerged in water, attached to surfaces above the bottom of the body of water. See also: benthos, plankton.
perisperm
The nutritive tissue in an angiosperm seed that surrounds the embryo, formed from the nucellus. See also: endosperm.
peristome
The fringe surrounding the mouth of the capsule upon removing the lid.
perithecium (pl. perithecia)
A small flask-shaped structure, containing asci.
perlite
A lightweight aggregate made from a volcanic glass that has been expanded by heat to form white, very lightweight kernels useful in opening cavities in soil to allow water and air to reach the roots; often used in potting soil.
permaculture
The maintaining of a permanent horticulture or agriculture by relying on renewable resources and compatibility with the local ecosystem.
permafrost
Frozen ground in arctic and subarctic areas which does not thaw in summer.
persistent
Lasting beyond maturity without being shed, as some leaves remaining through winter, etc.
personate
Describes a corolla which is bilabiate, and the throat closed by a prominent palate.
perula (pl. perulae, alt. perule)

One of the scales of a leaf bud. 2. A pouchlike portion of the perianth in certain orchids.
perulate
Describes leaf buds which are covered with scales.
petal
A division of the corolla; one of a circle of modified leaves immediately outside the reproductive organs, usually brightly colored.
petaliferous
Bearing petals.
petaline
Of or resembling petals.
petaloid

Colored and resembling a petal. 2. Additonal floral parts on ray florets having the form and appearance of smaller petals, e.g., in the Collarette dahlias.
petiolate
Having a petiole.
petiole (syn. leafstalk)
The stalk of a leaf that attaches to the stem.
petiolulate
Having a petiolule.
petiolule
The stalk of a leaflet.
phanerogam
A general name for flowering plants.
phaneropore
Superficial stoma.
phellem
A layer of usually suberized cells produced outwardly by a phellogen.
phelloderm
A layer of parenchyma produced inwardly by a phellogen.
phellogen (syn. cork cambium)
A secondary meristem that produces phellem and phelloderm in the periderm of a trunk or stem.
phenetic classification
The grouping of taxa by apparent similarities rather than evolutionary genetics.
phenology (adj. phenological)
The science of the relations between climate and periodic biological phenomena, e.g., the fruiting of plants or the color change of leaves.
phenotype
The morphological, physiological, behavioral, and other outwardly recognizable adaptations of an organism that develop through the interaction of genes and environment. See also: genotype.
phenotypic
Refers to a plant’s adaption to surrounding conditions, which are neither stable nor capable of being inherited (genotypic). Such visible changes occur especially where plants are grown in a wide variety of conditions, but will not carry over to different conditions, e.g., red leaves may occur in hot dry areas, but turn green when grown in normal conditions.
pheromones
Chemical substances produced by animals that attract and stimulate sexual partners of the same species.
photic zone
The upper layers of bodies of water into which sunlight penetrates sufficiently to influence the growth of plants and animals. See also: aphotic zone.
photohetrotroph
Describes an organism using light as a source of energy and organic materials as a carbon source.
photomania (adj. photomanic)
The response of an organism of seeking or growing toward light. See also: phototropism.
photomorphogenesis
The formation and differentiation of tissues and organs controlled by radiant energy, particularly light.
photoperiod
The duration of an organism’s daily exposure to light.
photoperiodic
Describes growth affected by exposure to light.
photophobia (adj. photophobic)
The dislike of light, as displayed by bugs that hide under rocks or bats which sleep all day and fly at night. See also: photomania, aphototropism.
photophosphorylation
The synthesis of ATP from ADP and phosphate that occurs in a plant using radiant energy absorbedduring photosynthesis.
photosynthesis
The manufacturing of sugar through the action of sunlight.
phototaxis
The movement of a body toward or away from a light source.
phototropism (adj. phototropic)
Growth or movement toward or away from a light source.
phreatophyte
A plant that can endure inundations of salt, such as cattails, Typha, which can live in estuaries, sieving out the saline molecules of seawater at a cellular level.
phyllary
One of the bracts under the flower head of a plant, especially in Compositae.
phylloclade (alt. phylloclad, syn. cladode, adj. phyllocladous)
A flattened, photosynthetic stem that performs the functions of a leaf, as occurs on some cacti. See also: cladophyll.
phyllocladia
Tiny life-like structures of some lichens.
phyllode
A flat expanded petiole that replaces the blade of a foliage leaf and fulfills the same functions. See also: cladophyll.
phyllodium (pl. phyllodia)
A somewhat dilated petiole having the form of and serving as a leaf blade.
phyllopodium
An outgrowth of the rhizome to which the frond is joined in some ferns.
phyllotaxy (alt. phyllotaxis)
The manner of leaf arrangement on a stem.
phylogenetic classification
The grouping of taxa by genealogical descent; evolution.
phylogeny (adj. phylogenetic)
The evolutionary development of a taxonomic group.
phylum
A major taxonomic grouping in the animal kigdom, ranking just below kingdom and above class. In the plant kingdom, it is usually replaced by the division.
physiographic climax
A pinnacle habitat controlled by the topography of the area; e.g., a forest growing on a north slope and a grassland on the south slope of the same ridge. See also: edaphic climax, biotic climax.
phytochemistry (adj. phytochemical)
The chemistry of plants, plant processes, and plant products.
phytochrome clocks
The coloring processes of plants that change the hues of ripening fruits and cause leaves to change colors with the shortened days of autumn.
phytogenesis
The evolutionary development of plants.
phytogenic (syn. phytogenous)
Having a plant origin, e.g., coal.
phytography
The science of plant description.
phytology (adj. phytological)
The study of plants.
phytopathogen
An agent-causing disease in plants.
phytoplankton
Small, often microscopic, aquatic plants.
phytotoxicity (n. phytotoxin, adj. phytotoxic)
Being poisonous to plants.
pick
A mattock type tool, which has one or both ends pointed.
pick pruning
The selection of individual branches to be cut; the opposite of shearing.
pier
A mass of masonry used as a support, breakwater, etc.
pilose
Hairy, especially with soft hairs.
pilosulous
Minutely pilose.
pinna (pl. pinnae)
One of the primary divisions of a pinnate or compoundly pinnate frond or leaf.
pinnate
Consisting of several leaflets arranged on each side of a common petiole or rachis on a compound leafor frond. 2. The feather vein pattern of simple leaves.
pinnately compound
With leaflets arranged in two rows along an axis.
pinnatifid (syn. pinnatipartite)
Divided in a pinnate manner, but with leaflets not entirely separate.
pinnatisect
Deeply cut, all the way to the axis.
pinnule
A secondary pinna; one of the pinnately disposed divisions of a pinna.
pip

The small seed of a fruit, like that of an apple or an orange. 2. An individual rootstock of lily of the valley or a similar plant.
pistil
The seed-bearing organ of the flower, consisting of the ovary, stigma, and style when present.
pistillate
Provided with pistils, and, in its more proper sense, without stamens.
pistillode
A sterile pistil, often rudimentary.
pitch
Lumps of resin found on the bark of trees.
pitcher-shaped
Campanulate, but with a distinct narrowing near the open end.
pith
The spongy or hollow center of twig or some stems.
pitted
Marked with small depressions or pits.
placenta
Any part of the interior of the ovary which bears ovules.
placentation
The arrangement of placentas within an ovary.
plankton
The floating or weakly swimming plants and animals occurring at any depth in bodies of water, often microscopic in size. See also: aeroplankton, edaphon.
plant
Any of the members of the kingdom Plantae typically lacking locomotive movement or obvious nervous or sensory organs and possessing cellulose cell walls and usually capable of photosynthesis.
plant labels

Plastic, wood, or metal stakes for gardens to indicate what seeds are planted where until they appear, and for varieties for evaluation. 2. Paper forms to include in drying plant samples, with formal printed forms as permanent labels on herbarium specimens. The minimum information required are the name of the collector, the location collected, the date collected, and hopefully, the correct identification of the specimen.
plant press (alt. field press)
A structure typically made of two ventilated frames 12 X 18 inches (30 X 45 cm.), within which plants may be arranged between sheets of driers and ventilators, tightly strapped together with press straps, in preparation of being added to a collection or herbarium.
plantation
A large farm of cultivated trees or plants, often of a single crop.
plantlet
A small plant, usually one produced vegetatively from a parent.
plasmodesma (alt. plasmodesm, pl. plasmodesmata)
One of the cytoplasmic strands that passes through openings in some plant cell walls and provide living bridges between cells.
plasmolysis
The shrinking of the cytoplasm away from the wall of a living cell due to the loss of water through osmosis.
plasticity
Flexibility; adaptability; capable of being repeatedly deformed without rupture.
plastid
Any of various cytoplasmic organelles of photosynthetic cells that serve in many cases as centers of special metabolic activities, e.g., chloroplasts.
plates
Flattened, rhizine-like structures on the lower surface of some lichens.
pleaching
A process whereby branches of woody plants are interwoven and plaited together to form an impassible hedge or very thick arbor. The only pruning done is to maintain a neat formal shape.
pleated
Describes a leaf creased along its length.
pleomorphic (n. pleomorphism)
Able to assume different forms.
pleurocarpous
Having the sporophyte lateral on a short lateral special branch. Pleurocarpous mosses can usually be recognized by the creeping habit.
plicae
Folds of a plicate leaf.
plicate
Folded into plaits, usually lengthwise.
plow
A device used to turn soil. On large farms several of these are mounted in a diagonal line, and referred to by the number of blades, e.g., a six-bottom plow. For compacted soil, chisels, four feet (1.3 m.) or longer, are used to open greater depth, and are called chisel plows.
plug
A core of grass sod or wildflower turf used in planting lawns and meadows.
plume
A feathery inflorescence.
plumose
Having fine hairs on each side, like the plume of a feather, as the pappus-bristles of some thistles.
plumule
The bud or growing point of the embryo.
plur- (alt. pluri-)
A prefix meaning many, e.g., plurilocular means many-celled.
plurilocular
Describes a sporangium or gametangium composed of many cells, each producing one zoospore or gamete.
pluriseriate
Many-ranked, as applied to leaves arranged in several rows along the stem.
pneumatophore
A specialized root in certain aquatic plants which performs respiratory functions.
pocket beach
The shore at a bay head where wave energy is lowest and settlings highest, often with fringe wetlands.
pod (syn. seedpod)
Any dry, several-seeded and dehiscent fruit.
podetium
A hollow upright structure.
polarity
The tendency for plants to develop from its poles, roots growing down, stems growing upward, making it essential to plant bulbs, etc, in the correct position.
pole pruner (syn. pruning stick, syn. pole saw)
A pruner on an extensible pole–usually having a curved saw at its end–which can be used to prune the upper parts of a tree or tall shrub.
pollarding
A process where tree tops are cut back severely each year to the same spots on the branches. This forces the growth of large knobby stubs from which long tender shoots grow each year.
pollen
The microspores of a seed plant contained in the anther, usually appearing as a fine dust.
pollen grain
A microspore of a seed plant.
pollen sac
The microsporangium of a seed plant where pollen is produced; the upper portion of the stamencontaining pollen grains; the anther.
pollinate (n. pollination)
To transfer pollen from the anther of a stamen to the stigma of a pistil, resulting in fertilization. This can occur either on a single plant (self-pollination) or between different plants.
pollinator
The agent, such as an insect or wind, which carries the pollen for fertilization.
polliniferous
Bearing pollen.
pollinium (pl. pollinia)
A mass of waxy pollen or of coherent pollen grains, as found in Asclepias and Orchidaceae.
polyad

A set of molecular states connected by vibratory resonances. 2. A patented name for the adsorptionprocess for the removal and destruction of Volatile organic compounds (VOC).
polybrid
A hybrid with more than two parental groups.
polyembryonic
Containing two or more embryos.
polyethylene
A plastic which allows the passage of gases but not of moisture.
polygamodioecious
Describes a plant group which has bisexual and male flowers on some plants, and bisexual and female flowers on others.
polygamous

Bearing male and female flowers on the same plant. 2. Having antheridia and archegonia disposed in various ways on the same plant.
polymorphic
Having more than two distinct morphological variants.
polypetalous
Having separate petals.
polyphyletic
Having members that originated, independently, from more than one evolutionary line. See also: monophyletic.
polyploidy (n. polyploid)
Having more than two sets of chromosomes in cells of the sporophyte or more than one set in cells of the gametophyte.
polysepalous
Having a calyx made up of separate sepals.
polystichous
Having leaves or other structures arranged in distinct rows.
polytrichous
Covered with many hairs.
polytypic
Containing more than one taxon of the next lower rank, e.g., a genus with more than one species. See also: monotypic.
pome
A kind of fleshy fruit, particularly an apple.
pompon
A small globular flower or flower head.
pond
A small, still body of water; an artificial body of water in a garden usually containing aquatic plants and fish.
pondweeds
A popular name for species of wetland plants that grow partially or wholly underwater.
pool
A small and relatively deep body of usually fresh water, either standing or as part of a stream. There is little wave action and often a reflective quality.
population
Plants of a species growing in a given place at a given time. Two similar species may mingle, e.g., Solidago canadensis and Solidago altissima, but the botanical term ‘population’ would include one species or the other, not both.
population density (syn. species density)
The concentration of individuals in relation to the space they occupy; how close individuals occur. See also: abundance, cover.
porate
Describes a pollen grain which has rounded apertures only. See also: colporate.
poricidal
A type of dehiscence in which the pollen is released through pores at the tip of the anther.
porose
Pierced with small holes or pores.
porrect
Spreading outward and forward.
Portland cement
A cement consisting predominantly of calcium silicates which reacts with water to form a hard mass.
positive drainage
Surface or underground pipes or trenches to remove excess water.
posterior
In an axillary flower, on the side nearest to the axis of inflorescence.
postreproductive
Refers to an organism which is past the age of breeding capacity.
potato fork
A fork of heavy construction with flat tines, for deep digging as is needed for potato harvesting.
potbound
The condition of a houseplant or outdoor container plant which has been left in the same pot or container for too long a period of time and the roots fill every niche of the pot or container. A hook or blade is required to cut through and separate the tangled matted outer roots when removed from the pot and before placing in the new container or the plant may never recover from this disastrous condition.
potential Hydrogen (abr. pH)
A log scale measurement of the acidity/alkalinity of a solution with 1 being extremely acidic, 10 being extremely alkaline, and 7 being neutral. Most plants prefer a soil within a certain range of pH.
potherb
A plant whose leaves, stems, or flowers are cooked or used as a seasoning.
pothole

A pond or pool utilized by waterfowl for nesting and raising young. 2. A hole worn in solid rock by the action of water, especially when a pebble repeatedly spins in the depression.
poultice
Moist, usually hot, mass of plant material; wrapped in a cloth and applied to the skin to bring about some desired action, such as bringing a boil to a head.
powdery mildew
A fungus forming a white powdery coating on leaves and stems.
power take off (abr. PTO)
Unit on a tractor to attach power tools.
Pozzolan cement
Volcanic rock powdered and used in making hydraulic cement.
pradines
Lopping tools.
praemorse
Appearing as if bitten off.
prairie
Grasslands, particularly that which formerly covered much of the central plains of North America, consisting of rich soil and a variety of grasses and forbs and generally covering a wide area.
precocious
Occurring early, as flowers appearing before the leaves; hysteranthous.
preemergent (alt. pre-emergent)
An herbicide, or combination fertilizer and herbicide, that is applied to a lawn early in the season, before the new growth has occurred.
prereproductive
Refers to a plant or animal which has not yet matured sexually.
prescribed burning
The intentional burning of plant material in an area. Sometimes used to consume underbrush and other fuel and thus prevent larger, more destructive fires. Also used to maintain a stable prairie or grasslandby aborting the process of succession to shrubs and trees.
press straps (syn. web straps)
Bands or ropes to hold the plant press tightly together, traditionally made of web and buckles, but now often with Velcro.
prevernal
Appearing in late winter or very early spring, as do crocus flowers.
prickle
A small, sharp outgrowth involving only the outer epidermal layer, usually more slender than a thorn. This is the correct term for rose thorns.
primary forest (syn. natural forest)
A tract of trees with minimal impact from humans.
primocane
The floricane, but in the first year of growth before it is capable of flowering.
primordial utricle
The layer of protoplasm lying next to the cell wall, which is very often conspicuous when dried and shrunken.
primordium
A tissue or organ in the early stages when it has undergone differentiation changes but has not yet matured enough to emerge.
priority
The rule of nomenclature whereby the first published of two validly published names for the same entity is given approval as the accepted name.
prismatic
Of the shape of a prism, angular, with flat sides, and of nearly uniform size throughout.
pro hybrida (abr. pro hybr.)
Means “as a hybrid.” Used in citations to show that a plant now known as a species started as a hybrid.
pro parte (abr. p.p.)
Means “in part”. Used in citations to show that a taxon as used by one author shows only a portion of what was intended by the original author.
pro specie (abr. pro sp.)
Means “as a species.” Used in citations to show that a plant which had been accepted as a species is now designated as a hybrid.
proboscis
Beak.
procumbent
Lying on the ground or trailing but without rooting at the nodes.
profundal zone
The area of deep water at the bottom of a lake below effective penetration of light. See also: abyssal.
projecting
Refers to stamens, styles, and stigmas which exert beyond the petals of a flower.
prokaryote (alt. procaryote, adj. prokaryotic, adj. procaryotic)
An organism composed of one or more cells lacking a visibly evident nuclei, including the viruses, bacteria, and cyanobacteria. See also: eukaryote.
proliferous (syn. proliferating)

Freely producing offshoots, bulblets, or plantlets. 2. In mosses, bearing young shoots from the antheridial or archegonial cluster of leaves.
prominent
Standing out from the surface, clearly visible and touchable, as are veins and other surface features.
prominulous
Slightly raised above the adjoining tissue.
propagate (syn. propagation)
To produce new plants, either by vegetative means involving the rooting or grafting of pieces of a plant, or by sowing seeds.
prophyll

The bracteole under a single flower or pedicel. 2. The showy first bract borne on the peduncle and, occasionally, on the inflorescence branches of some monocots.
prosenchymatous
Cells with pointed ends dovetailing into each other.
prostrate
Lying flat upon the ground.
protandrous
Refers to a flower where the shedding of the pollen occurs before the stigma is receptive. See also: protogynous.
protected area
A legally restricted land or water area under public or private ownership, which has managed to achieve specific conservation objectives.
proteinaceous
Of, relating to, resembling, or being protein.
proterogynous
Having the stigma ripe for the pollen before the maturity of the anthers of the same flower.
prothallium (pl. prothallia)
The minute scale-like growth from the spore of a fern.
prothallus
A cellular, usually flat and thallus-like growth, resulting from the germination of a spore, upon which are developed sexual organs or new plants. The gametophyte stage of ferns.
protogynous (syn. proterogynous)
Refers to a flower where the shedding of the pollen occurs after the stigma has ceased to be receptive. See also: protandrous.
protologue
Refers to all parts of botanical publication, including, but not limited to, diagnosis, description, synonymy, citation of specimens, illustrations, etc., expressed as data, information, and knowledge.
protonema
The green, branched, alga-like threads produced from the spore and often persistent during the lifetime of the plant produced from it.
protoplasm
The living contents of a cell.
protoxylem
The first formed xylem.
proximal
Toward the base.
proximate
Close together.
pruina
A woolly white covering of the upper cortex of some lichens.
pruinose
Having the surface obscured by a rather thick, bluish or grayish, wax-like coating.
prune
To cut back, for the purposes of shaping a plant, encouraging new growth, or controlling size.
pruning saw
A serrated blade tool for cutting small limbs; may be styled so the blade folds into the handle, or fit into a sheath.
pruning shears
Specialized scissors to cut plants back.
pseudanthium (pl. pseudanthia)
An inflorescence which looks like a simple flower, but is made of more than one axis with a number of flowers.
pseudobulb (adj. pseudobulbous)
A swelling at the base of a stem in which epiphytic orchids store nutrients and water.
pseudocarp (syn. accessory fruit, syn. false fruit)
A fruit, such as the strawberry or pear, that develops not only from the ripened ovary, or ovaries, but from nonovarian tissue as well.
pseudocyphellae
Pores found in the upper or lower cortex of lichens.
pseudolamina
A false lamina, one usually formed by the lateral expansion of a rachis.
pseudoparenchyma
Filamentous tissue compacted to resemble parenchyma.
pseudopodetium
An upright, fruticose thallus of some lichens.
pseudopodium
A leafless branch resembling a seta and often bearing gemmae.
pseudopods
Literally, false feet; a temporary protrusion of the protoplasm of a cell for the purpose of moving or feeding; foot-like organs.
pseudostem
An erect aerial growth which appears to be a stem with leaves, but is actually packed or overlapping sheaths and stalks of essentially basal leaves.
pseudoterminal

Describes a bud which is situated at what appears to be the apex of a branch, but is in fact located in an axillary position beside a leaf scar. 2. Refers to a bud which replaces an injured terminal bud which has died.
pteridology (n. pteridologist, adj. pteridological)
The study of ferns.
pteridophyte
Any of the nonflowering vascular plants of the division Pteridophyta having roots, stems, and leaves; a fern or fern ally.
Pterophyta
A major taxonomic division containing the ferns, showing clear alternation of generations with a dominant vascular sporophyte which begins with the very reduced gametophyte.
puberulose (alt. puberulent, alt. puberulous)
Minutely pubescent.
pubescence
A covering of soft, short hairs.
pubescent
Covered with hairs, especially if short, soft, and down-like.
puddle

A very small pool of water, often transitory, appearing during heavy rain and disappearing shortly afterward. 2. To knead or work soil while wet, making it impervious to water. 3. To work while wet, as in flooded rice fields.
Pulaski clearing axe (alt. Pulaski)
A tool used mostly by firefighters that is a combination of axe and grub hoe.
pulp

The soft, succulent part of a fruit, usually composed of mesocarp. 2. The pith of a stem.
pulpwood
The wood of trees like aspen, Populus, and spruce, Picea, which is easily ground into a soft mass for the production of paper.
pulverulent
Powdered; appearing as if covered by minute grains of dust.
pulvinate
Like a cushion.
pulvinus
A swelling at the base of the stalk of a leaf or leaflet.
punctate
Dotted with depressions or with translucent internal glands or colored dots.
puncticulate
Minutely punctate.
pungent
Terminating in a rigid sharp point; acrid to smell or taste.
purgatives
Medicines which empty the intestines.
pustular
Like a blister.
pustulate
Covered with blisters.
pustule
A surface eruption such as a pimple or blister.
putamen
The shell of a nut; the bony part of a stone fruit.
PVC sleeves
A plastic pipe used for moving water that acts as a conduit.
pycnidium (pl. pycnidia)
A rounded or flask-shaped asexual fruiting body containing spores found in certain fungi.
pyramidal
Conical with flat angular sides.
pyrene
The stone of a succulent fruit made up of the endocarp and the seed. See also: drupe.
pyrethrin
The pyrethrins are a class of organic compounds normally derived from Chrysanthemum cinerariifolium that have potent insecticidal activity by targeting the nervous systems of insects. Pyrethrin is synthetically made by industrial methods, but it also naturally occurs in chrysanthemum flowers, thus is often considered an organic insecticide.
pyriform
Pear-shaped.
pyxis
A capsule which opens in a circumscissile manner.
quadrate
Describes cells that are square or nearly so.
quadrifid
Divided into four parts.
quadrilateral
With four sides.
quadrinary combination
Binomial plus two more taxon names such as subspecies and variety.
quadripinnate
Four times pinnate.
quarantine
A legal ban on the export or import of certain noxious weeds or insects that may be attached to the plants.
quartered
Having the petals formed into a number of sections, usually four but sometimes three or five.
quinate
Having five leaflets growing from the same point of attachment.
quincunx (alt. quincunce, adj. quincuncial, adj. quincuntial)
The arrangement of five objects, such as trees, with one on each of four corners and one in the center.
raceme
A simple inflorescence of pediceled flowers upon a common more or less elongated axis.
racemiform
Refers to an inflorescence with the outward appearance of a raceme.
racemose
Resembling a raceme.
rachilla
The axis of a spikelet, particularly in grasses.
rachis

The main stalk of a flower cluster or the main leafstalk of a compound leaf. 2. In ferns, the continuation of the stipe through a compound frond.
radial
In flowers, one with the symmetry of a symmetry wheel.
radiate
Spreading from or arranged around a common center; bearing ray flowers.
radical
Belonging to or proceeding from the root or base of the stem near the ground.
radicle
The portion of the embryo below the cotyledons that will form the roots, more properly called the caudicle.
radicles (alt. rhizoid)
Rootlets springing from the sides and base of the stem.
radius (pl. radii)

The outermost flowers in a daisy-like head, Compositae; ray flowers as opposed to disc flowers. 2. The distance from the center of a circle to the outer edge, as from the heart of a tree to the bark.
raffia (alt. raphia)
A fiber-like material used for tying grafts and other horticultural purposes.
raised beach
An old beach terrace at an elevation above the present shoreline, representing a formerly higher lake level.
raised bed (syn. raised garden)
A bed or garden created by adding soil on top of that which is already present. This is usually done in cases where the underlying soil is particularly poor in nutrients or has inadequate drainage. Wooden or stone walls are often used to keep the added soil in place.
ramble
To grow freely, often over another plant or structure.
rameal
Belonging to a branch.
ramentaceous
Refers to stems or leaves which have small, loose, brownish scales.
ramet

An individual member of a clone. 2. An underground tree system giving rise to large suckering colonies, like big-tooth aspen trees, Populus grandentata.
ramification
Branching.
ramiflorous
Bearing flowers directly from large branches and leafless twigs, but not on the trunk.
ramiform
With branches; branch-like.
ramose
Having many branches.
ramuli
Minute branchlets.
ranch (alt. ranche, rancho)

In the united States and Canada, a farm that raises livestock, fruits, and vegetables for use of the family, or crops for feeding animals; sometimes used as a secondary income, e.g., a cattle ranch. 2. The buildings occupied or the persons on the estate (ranchers).
range of variation
The extremes of possibilities which may occur within a species and still remain a member of that species, including such things as leaf shape, leaf margins, flower color, height, etc.
rank

A vertical row, as of leaves. When you sight along the length of a branch from the tip end, if it appears there are two rows of leaves, either opposite or alternate, the branch is 2-ranked; if three rows, it is 3-ranked, etc. 2. In taxonomy, the position of a taxon in the hierarchy, e.g., species, genus, family, etc.
raphe
The part of the funiculus that is fused along the side of the ovule.
raphides
Needle-like crystals of calcium oxalate that occur in bundles in some plant cells.
ratchet tool
One which uses mechanical means to multiply cutting or turning power, reducing cramping of hands and forearms.
ravine
A depression worn by running water, larger than a gully and smaller than a valley.
ray
The branch of an umbel; the marginal flowers of an inflorescence when distinct from the disk.
ray floret
One of the broad, conspicuously colored florets of a compound flower, the structure of which suggests a single petal of an ordinary flower. These form the radiating border in the open-centered type dahlias, or massed together, the flower head in fully double types. See also: disc floret.
ray flower
The bilaterally symmetrical flowers around the edge of the head in many members of Compositae; each ray flower resembles a single petal.
rays
Radial strands of cells in wood and phloem.
recalcitrant seed
Seed that does not survive drying and freezing.
receptacle

The more or less expanded or produced portion of an axis which bears the organs of a flower (the torus) or the collected flowers of a head, and in roses, enfolds the developing ovaries to form a hip. 2. Any similar structure in cryptogams.
receptacular
Carried on the receptacle; pertaining to the receptacle.
recession
A drop in the water line of a body of water, with a corresponding exposure of the bottom.
reclinate
Bent or turned downward.
reclining
The lower portion somewhat flattened along the ground but the upper parts curving upward.
recurved
Curved downward or backward; with ray florets curved backward along their length toward the stem.
reeds
Tall thin wetland plants of the genera Phragmites and Sparganium; a term often incorrectly applied to bulrushes.
reel mower
A grass-cutting tool on which helical blades spin against a flat surface, generally hand powered but may be mechanized for a more precise cut than rotary mowers.
reference
A citation to a source of information, such as a written document.
reflexed
Abruptly bent or turned downward.
refrigerants
Herbal preparations which cool the body.
regular
Uniform in shape or structure.
regular flower
Generally symmetrical and uniform in the number of its parts.
rehabilitation
The process of improving specific ecosystem factors in a degraded habitat.
rejuvenation pruning
The practice of cutting all the main stems of a shrub back to within half-inch of the ground during winter dormancy.
relict (alt. relic)
A species or other group within a community that is representative of an earlier stage of development or of a different set of conditions.
relief
The difference in elevation between the highest point and the lowest point of land in a specified region.
remontant
Able to rebloom one or more times during a single growing season.
remote
Distant, at the farthest end.
renewal pruning
The practice of cutting all the main stems of a shrub back to within half-inch of the ground during winter dormancy.
reniform
Kidney-shaped.
repand
With a slightly uneven and somewhat sinuate margin.
repellent
A substance used to keep insects or animals from an area.
repent (alt. repen, syn. reptant)
Creeping; prostrate and rooting at the nodes.
replicates
Duplicate specimens.
replum
The hardened framework of the placenta which remains when the outer walls or valves of a silique fallaway; the septum which holds the seeds.
resin
A plant secretion, often aromatic, that is insoluble in water but soluble in ether or alcohol.
resin dot
Tiny circular or globular yellow spots, usually not obvious except under magnification.
resiniferous
Producing resin.
resinous
Coated with a sticky gum or resin.
restoration
The return of an ecosystem or habitat to its original structure, natural complement of species, and natural functions.
resupinate
Turned upside down.
resupine
Refers to a flower or leaf on which the stalk twists a full half turn, 180 degrees, so that the organ appears to be upside down.
retardants
Admixtures that increase the setting time of cement by slowing down hydration.
retentive sepals
Sepals that remain attached to the apex of the receptacle after it has ripened into a hip.
reticulate
In the form of network; netveined.
reticulation
The entire network of reticulate veins, ribs, coloring and/or fibers.
reticulum (pl. reticula)
A mesh-like network.
retinaculum
The structure by which another structure is attached to a plant.
retrorse
Directed back or downward.
retting
Soaking plant tissue so as to induce bacterial growth that will aid in the separating of the fibers.
retuse
With a shallow notch at a rounded apex.
reversion
When a genetically differentiated plant or sport reverts to normal growth. For instance, variegatedleafs turn green.
revolute
Rolled backward from the margins or apex; describing a ray floret with margins rolled backward along their longitudinal axis. See also: fully revolute.
rhachilla
A secondary axis; specifically, in the grasses and sedges, the floral axis as opposed to that of the spike or spikelet.
rhachis
The axis of a spike or of a compound leaf.
rhape
The ridge or adnate funicle which in an anatropous ovule connects the two ends.
rhaphides
Needle-shaped crystals often occurring in bundles within the cells of certain plants.
rhizines
Strands of hyphae found on the lower surface of many foliose lichens.
rhizobia
Bacteria in a symbiotic relationship with leguminous plants that results in nitrogen fixation.
rhizomatous
Having or appearing to have rhizomes.
rhizome (syn. rootstock, syn. understock, adj. rhizomatous)
Any prostrate or subterranean stem, usually rooting at the nodes and becoming erect at the apex.
rhizome chaff
Small pieces of rhizomes which are spread upon the ground and covered with soil so they can take rootand form new plants.
rhizome cutting
A piece cut from a rhizome which can take root and form a new plant.
rhizosphere
The area that surrounds and is influenced by a plant’s roots.
rhombic
Diamond-shaped.
rhomboid
Refers to leaves, tepals, etc., which are diamond-shaped, with the base and tip having acute angles and the sides having obtuse angles.
rib
A primary or prominent vein of a leaf.
ribbed
With one or more prominent veins or nerves.
ribs
Geologic ridges running parallel, as often occurs along a mountain side.
ridged
Angular, with lengthwise lines.
rind

The bark of a tree outside the cambium layer. 2. The epicarp of a citrus fruit, e.g., an orange or a lemon.
ringed
Having narrow encircling stipule scars around leaf scars.
ringent
Gaping, as the mouth of an open bilabiate corolla.
riparian
Of, or relating to, rivers or streams.
ripening

The maturing of a fruit as developmental changes prepare it for seed dispersal. 2. The hardening of wood, especially of soft twigs, important in taking cuttings for propagation.
risers
The up and down sections of stair steps; necessary for landscape plans.
riverine
Refers to a river.
rivermouth
The place where a river empties into another body of water.
rock garden
A garden laid out among rocks and adapted for the growth of particular kinds of plants, such as alpines.
rolled
Curled under; revolute.
root
The underground part of a plant that serves to anchor it and supplies it with nourishment.
root cutting
A cutting taken from the root of a parent plant for the purpose of propagation.
root knot
A disease of the roots characterized by a swelling and caused by nematodes.
root pruning

Pruning to stimulate the growth of new roots within a parameter in preparation for transplanting. 2. Trimming and/or scoring the outer layer of roots of a plant that has become root bound. 3. The act of removing a portion of a plant’s roots to keep top growth in check.
root zone
The area immediately surrounding the roots and from which a plant takes moisture and nourishment.
rootball (alt. root ball)
The root stock and surrounding soil together; in nurseries and during transport, these are held together by burlap or other wrapping.
rootbound (alt. root bound)
The condition of a houseplant or outdoor container plant which has outgrown its container, with the roots filling every niche of the container. Eventually, the roots will themselves strangle the plant by constricting the flow of nutrients. A hook or blade is required to cut through and separate the tangled matted outer roots when removed from the pot and before placing in the new container.
rootlet
A small, secondary root.
rootstock (syn. rhizome, syn. understock)

A rhizome. 2. The root system and lower portion of a woody plant to which a graft of a more desirable plant is attached.
rootworm
The larvae of a beetle of the genus Diabrotica that feed on the roots of various plants, particularly corn.
roridulous (alt. roridulate)
Dewy; covered with waxy plates which appear to be dew drops.
rose dethorner
Florist tool to remove the thorns from roses and other flowers.
rose hip
A smooth, rounded, fruit-like structure consisting of the cup-like calyx enclosing seed-like fruits.
rosette
A cluster of leaves which grows in a circular overlapping pattern–somewhat like the petals of a rose–arising basally from a crown, as on a dandelion, or apically from an axis, as on many palms.
rostellate
Having a short beak.
rostellum
A little beak: a slender extension from the upper edge of the stigma in orchids.
rosula
A rosette.
rosulate
In the form of a rosette.
rot
Any of various diseases causing the breakdown and rotting of tissue.
rotary edger and trimmer
A lawn tool which has rotating spikes on one edge to trim grass back from drives and walks.
rotary mower
The most common type of grass-cutting tool, usually powered by gasoline or electricity.
rotate
Describes a corolla that is wheel-shaped flat and circular in outline.
rotation grazing
Alternating between two or more pastures. See also: cell system pasturing, deferred grazing.
rotund
Rounded.
rough
Papillose
rubefacients
Herbal medicines which open the small blood vessels of the skin to increase its heat and redness.
ruderal
Growing in poor land or waste places.
rudimentary
Partially developed.
rufescent (alt. rufous)
Reddish-brown.
rugulose
Covered with minute wrinkles.
ruminate
Having a surface which is coarsely wrinkled, appearing as though chewed by a cow and then spit out, but not with the margins appearing gnawed as in erose margins.
runcinate
Sharply incised, with the segments directed backward.
runner
A filiform or very slender stolon. A prostrate branch that roots at its joints.
runoff (alt. run-off)
Water which originated on top of the land, such as rain, and collected beyond the ability of the soil to absorb it, so it flows away.
rush (pl. rushes)
Any of various monocotyledonous, grass-like, often tufted marsh plants of the family Juncaceae, having cylindrical often hollow stems.
rust
A large fungus group infecting such plants as roses and snapdragons. Round spots occur on the leaves, filled with yellow or reddish spores.
rusticate (n. rustication)
To build or face with usually rough-surfaced masonry.
saccate
Sac-shaped.
sagittate
Shaped like an arrow-head, the basal lobes directed downward.
salinity
Refers to an excess of salts in the soil which causes leaves to scorch and turn yellow and does great harm to many plants.
salver-shaped
Describes a corolla that has a slender tube abruptly expanded into a flat limb.
salverform
Having a slender tube which expands abruptly.
samara
A dry, one-seeded, indehiscent winged fruit; a key.
samaroid
Resembles a samara, a winged fruit.
sandbar (alt. sand bar)
A temporary ridge of sand formed by currents in a body of water.
saponin
A glycoside compound of plants which has a foaming or soap-like action.
saprophyte
A plant lacking chlorophyll and living on dead organic matter.
saprophytic
Feeding from dissolved or decayed organic material, as a saprophyte.
sapwood
The more recently produced, outer layer of wood tissue of a tree, usually lighter in color than the heartwood.
sarmentose
With long slender stolons.
saturation cratering
Ancient land around a volcanic area, filled with old and new craters from repeated eruptions.
savanna (alt. savannah)
A flat grassland of tropical and subtropical regions usually having distinct periods of dry and wet weather.
saxatile (alt. saxicolous, alt. saxicoline)
Growing among or upon rocks.
scaberulous
Slightly scabrous.
scabrid
Minutely rough.
scabridulous
Slightly rough.
scabrous (syn. scabrid)
Rough to the touch.
scaffold branches
The main side branches of a tree, especially a fruit tree.
scalariform
Having a ladder-like pattern.
scale

A thin, membrane-like covering of the bud or twig base. A fine, grain-like surface material. 2. Sap-feeding insects that live under shell-like scales on stems, bark, and leaves.
scale leaves
Specialized leaves, including those covering buds or composing bulbs.
scales
The chaff on the stems of ferns.
scaly
Having tiny scales attached at only one end.
scandent
Climbing, but not self-supporting.
scape
A peduncle rising from the ground, naked or without proper foliage, e.g., a dandelion; the leafless stemof a flower.
scapose
Bearing or resembling a scape.
scarify (n. scarification)
To scar or nick the seed coat to enhance germination.
scariose
Scarious, having thin, dry, shriveled tissues.
scarious
Thin, dry, and membranaceous, not green.
scarred
Having marks where organs have fallen off, such as leaf scars on stems.
schizocarp
A fruit that splits between carpels into one-seeded portions or lobes.
scientific name (syn. botanical name, syn. technical name)
The name applied to a plant, animal, or other organism, according to the Codes of Nomenclature. Although in some cases a scientific name is changed; it is almost always a much more reliable manner of identifying a plant than the often variable common name. See also: binomial.
scion
A cutting from the upper portion of a plant that is grafted onto the rootstock of another plant, usually a related species.
scion rooting
Covering a low graft with soil so that the plant develops roots directly from both the rootstock and the scion.
scion wood
Shoots from which graft scions are cut.
sclereid
A variably shaped sclerenchymatous cell of a higher plant.
sclerenchyma (adj. sclerenchymatous)
Tissue composed of cells with thickened and hardened walls.
scleromorph
A plant with leaves and/or stems that are hard, usually having a thick cuticle. See also: xeromorph.
sclerophyll
A woody plant with small, leathery, evergreen leaves and generally found in hot, dry climates. See also: chaparral, fynbos.
sclerophyllous
Having leaves stiffened by sclerenchyma.
sclerotium
A dense mass of filaments comprising the dormant phase of certain fungi.
scoop shovel
A shovel with a very large blade, often made of thin light-weight metal or plastic, used for moving large amounts of crops such as ear corn, or clearing water.
scorch
The drying and browning of leaf margins usually caused by unfavorable environmental conditions.
scorpioid
Describes an inflorescence that is circinately coiled while in bud.
scrambler
A plant with long stems, often with thorns to help it support itself as it climbs through shrubs.
scree
Fragmented rocks usually found at the base of a cliff or boulder where the action of wind and water has broken the particles loose. Scree may be used in gardens as a mulch.
screen
A single plant or grouping of plants used to bar certain parts of the landscape from view.
scuffle hoe
A medium-weight hoe with the blade fastened to the handle in the middle of the horizontal blade, allowing it to be pushed and pulled, cutting weeds on each direction.
scutate (alt. scutellate)
Platter-shaped, with a flat center and raised margins, as a water lily leaf, Nymphaea lotus.
scutelliform
Shaped like a small shield.
scythe
A long-handled blade for making grass into hay; the term is sometimes used for the blade without the handle. See also: snath.
seawater
Ocean water with a high salt content.
secateur
Hand-held pruning shears.
secondary species
The species subordinate to the dominant species, like dogwood, Cornus, in an oak, Quercus, forest.
sectio (alt. section)
The category of supplemental taxa at a rank between subgenus and series.
secund
Twisted or turned to one side.
sedatives
Medicines which can slow certain bodily functions and promote sleep.
sedge
Any of the grass-like, monocotyledonous plants of the family Cyperaceae, having achenes and solidstems wich are triangular in cross-section.
seed
The ripened ovule, consisting of the embryo and its proper coats.
seed coat
The outer protective covering of a seed.
seed drill

A shallow trench in which to sow seeds. 2. An implement for planting seeds.
seed fern
Extinct fern-like plants that bore seeds on the frond.
seed leaf
A cotyledon.
seed lot
Seeds of a particular crop gathered at one time and likely to have similar germination rates and other characteristics.
seedbank
A facility designed for the ex situ conservation of individual species and varieties through controlled seed storage.
seedbed
An area cultivated for planting seeds.
seedhead
The dry fruit containing rip seeds.
seedling
A young plant grown from a seed.
segment

One of the parts of a leaf or other like organ that is cleft or divided.
segregate

A new genus culled from an established one for taxonomic reasons. 2. A hybrid which exhibits the genetics of one of the parents more clearly than those of the other parent.
seiche
A rapid and sometimes violent fluctuation in water level within a landlocked body of water, usually due to sudden changes in barometric pressure, the ripples lasting from minutes to several hours. See also: tsunami.
selection
A distinct form of a plant, sexual or asexual in origin, selected and propagated for monetary reasons or beauty, and named as a cultivar.
selective felling (syn. selective cutting)
When only choice trees are cut in a forest.
self
Having a single pure color with no markings.
self-clinging
Refers to plants which climb rocks or structures without the support of other plants, perhaps by suckers or tendrils.
self-fertile
Able to be pollinated by its own flowers.
self-pollination
The transfer of pollen from the anther of a flower to the stigma of the same flower, or to different flowers on the same plant.
self-seeding
Produces offspring from seed without intervention.
self-sowing
Produces offspring from seed without intervention.
semelparity
Refers to an organism that has a single reproductive season and then dies. See also: iteroparity.
semi-
Prefix meaning half or partly.
semidouble (alt. semi-double)
Having more than the usual number of petals but with at least some pollen-producing stamens.
semievergreen (alt. semievergreen)
Retaining at least some green foliage well into winter, or shedding leaves only in cold climates.
semihardwood cutting
A cutting taken from a stem that has just begun to develop woody tissue, for the purpose of propagation.
semihardy
Questionably hardy in a given temperature zone; susceptible to damage by extreme cold or heat.
semirotate
Saucer-shaped, almost flat.
semiterete
Refers to a cylinder which is rounded on one side, but flat on the other, as some leaf petioles.
senescence
The process of aging in a plant or plant part (as a leaf) from full maturity to death.
senescent
Dying of tissues following maturity.
sensu
Meaning “in the sense of” in taxonomy.
sepal
A division of a calyx; one of the outermost circles of modified leaves surrounding the reproductive organs.
sepaline
Refers to sepals.
sepaloid
Like a sepal.
sept
A small dent or cavity.
septate
Divided by partitions.
septicidal
Describes a capsule that is dehiscing through the partitions and between the cells.
septifragal
Describes the dehiscence of a fruit where the valves or backs of the carpels break away leaving the septa intact.
septum (pl. septa)
Any kind of partition.
sere
The complete cycle of changes from the original state to the climax condition, e.g., oak-maple forest, forest destroyed by fire, meadow, brushland, young mixed conifer-deciduous forest, oak-maple forest.
seriate
In a whorl or row, generally preceded by a number or numerical prefix, as triseriate meaning in three rows.
sericeous
Bearing fine, usually straight, appressed, hairs.
series
A group of similar, but not identical, plants.
serotinal
Appearing in the autumn or late summer.
serrate
Having sharp teeth pointing forward.
serrulate
Finely serrate.
sessile
Without footstalk of any kind.
set

The development of fruit, and or seed, following pollination; to set seed. 2. To transplant as seedlings. 3. To apply as a graft. 4. A young bulb, tuber, or other type of vegetative propagule ready for planting. 5. Transformation of cement paste or concrete from a fluid-like consistency to a stiff mass.
set seed (alt. go to seed)
To produce seeds after flowering. In some cases a gardener wishes to prevent this in an effort to prolong flowering, or continue harvesting the foliage of an herb.
seta

A bristle. 2. In mosses, the stalk holding the spore capsule.
setaceous
Bristle-like.
setiform
Bristle-shaped.
setose
Beset with bristles.
setulose
Having minute bristles.
sextant
An instrument used mostly at sea, where there are no bench marks, to determine location by latitude and longitude.
shade (adj. shady)
Light diminished by some blocking feature. See also: light shade.
shade tree
A larger tree planted in a location where the canopy provides shade at ground level.
sharp sand
Coarse sand used to loosen heavy soil and in construction, as opposed to the more finer grained play sand.
sharp-pointed
Acute, mucronate, pungent.
sheath
A tubular envelope, as the lower part of the leaf in grasses.
sheathing
Enclosing as by a sheath.
shoot
A young, actively growing twig or stem.
short-lived
Describes a plant, usually perennial, where the individuals live for only a few seasons.
shovel

A long-handled scoop-like implement used for digging, lifting, and throwing, etc. 2. The quantity contained in one scoop.
shrub
A woody perennial, smaller than a tree, usually with several stems.
shrublet
A dwarf woody plant.
sialagogues
Herbal medicines that can increase the flow of saliva.
sickle

A short-handled hand tool similar to a scythe but smaller and curved into a semicircle, used to cut heavier crops like corn, Zea mays.
sickle bar
A mechanical cutting tool that fastens onto a tractor’s power source.
sigmoid
Curved like the letter S.
siliceous
Containing silica.
silicle (alt. silicula, alt. silicule)
A short silique, no more than twice as long as it is broad.
silique (syn. siliqua)
A dry, dehiscent, elongated fruit formed from a superior ovary of two carpels, with two parietalplacentas and divided into two loculi by a false septum between the placentas, occurring in plants of the family Cruciferae.
silky
Covered with close-pressed soft and straight pubescence.
silt
Fine-grained sediment usually 1/20 millimeter or less in diameter, finer than sand particles and coarser then those composing clay.
silt load
The particles finer than sand which are carried in a suspended state in moving water. See also: bed load.
silviculture
The cultivation and care of forest trees.
simple
Of one piece; not compound.
simple fruit
A fruit developed from a single ovary.
simple leaf
A leaf with an undivided blade.
single

A single flower. 2. Describes varieties having only one bloom per stem.
single flower
A plant containing the normal number of petals. Double flowers often have multiples of the normal number: 6 or 9 for three-petal flowers, 8 or 12 for four-petals flowers, etc.
single-worked
Grafted only once so that the grafted plant consists of rootstock and scion.
sinuate
With the outline of the margin strongly wavy.
sinuous
Wavy like the path of a snake.
sinus
The cleft or recess between two lobes.
skeletonized
Refers to a leaf which has only ribs and veins left, the soft tissue having been consumed by insects or bacteria.
skeletonizer
Any of various lepidopterous larvae that eat the parenchyma of leaves reducing them to a skeleton of veins.
skylight
The diffuse and reflected light of the sky. See also: direct sunlight.
sludge
Precipitate of solids from water or sewage treatment, which may or may not contain harmful compounds; generally composted for a year or two before applying as organic fertilizer.
slump test
A test used to determine workability of cement or concrete.
small fruits
Those species which produce edible fruits, but do not grow on trees, e.g., strawberries and blueberries.
smoke jumper
A trained firefighter who is dropped from an airplane in the path of a wildfire to attempt to control the blaze.
smooth
Without roughness or pubescence.
snag

Part of the rootstock left during a graft to hold the scion in place. 2. Removal of the projection when the scion no longer needs it. 3. A rough projection on a woody plant which needs to be removed.
snath
The handle of a scythe.
snow fencing
Wooden or plastic lath joined by wire into long rolls, then erected 100 feet (33 m.) or so on the windward side of a road or barnyard. The lath breaks the velocity of the wind allowing snow particles to drop before reaching the road. It may also be used as a shade cover for plants.
sobole
A shoot, especially from the ground.
soboliferous
Bearing soboles.
social dominance
The control of behavior of those animals lower in the hierarchy, by aggression, intelligence, or other factors of the alpha animals.
social facilitation
The effect of the closeness of one organism upon the conduct of another one.
sod coring tool
A lawn tool which removes spikes of soil, about one half inch (1 cm) in diameter, to allow air and water to penentrate. See also: aerator.
sod planter
A lawn tool which cuts a hole of about 2 inches (5 cm.), releases the core with a hand or foot system, picks up a new sod block and inserts it.
sodic
Of, relating to, or containing sodium.
softwood
The immature stems of woody plants. Also, term applied to conifers as opposed to the broad-leavedtrees, or hardwoods.
softwood cutting
A cutting taken from a green or immature stem of a woody plant, for the purpose of propagation.
soil

The top layer of the earth’s surface, consisting primarily of clay, sand, silt, and organic matter. 2. Any natural or synthetic substance or medium in which plants may take root and grow.
soil amendment
Substance added to soil to increase its nutritive value, friability, moisture retention, or some other aspect.
soil cuber (alt. soil blocker)
Hand-held device into which moist soil is packed, then released in the form of a cube or block in which a single seed is planted.
soil fixation
The conversion of a soluble substance such as phosphorus from the exchangeable form useful to plants, to a relatively insoluble form.
soil sampler
A hollow tube with a T handle, which is pressed into the soil to get a core of the horizon.
soilless mix
A potting soil that uses sand, bark, and/or other ingredients, but not any actual soil.
solid
Smoothly pithy, the twig center neither chambered nor hollow.
solitary
Borne singly or alone; not in clusters.
solutio (abr. sol)
Latin term for a solution.
somatic
Belonging to the vegetative part of a plant as opposed to the reproductive.
soralium (pl. soralia)
A clump of soredia.
soredium (pl. soredia)
A microscopic clump of agal cells surrounded by the hyphae of fungi and erupting at the surface of the thallus.
sorus (pl. sori)
An assemblage of sporangia; a fruit dot.
spacing
Measured distance between the center of one plant and the center of the next closest ones.
spade
A digging implement, foot powered, usually of heavier construction than a shovel and often with a flat end.
spadix
A flower spike with a fleshy axis, usually enclosed in a spathe.
spartoid
Rush-like, but sharp and rigid.
spasmolytic
Describes an herbal preparation that can ease cramps or spasms.
spathaceous
Resembling or having a spathe.
spathe
A sheathing bract or pair of bracts partly enclosing an inflorescence and especially a spadix on the same axis.
spatheole
A small or secondary spathe.
spathiform
Shaped like a spathe.
spatulate (alt. spathulate)
Gradually narrowing downward from a rounded summit; spoon-shaped.
species (abr. sp., pl. spp.)
A fundamental category of taxonomic classification that ranks below a genus and above subspecies; a population or series of populations whose individuals have the potential to freely breed with one another and that is discontinuous in variation from other populations or series of populations. The plural abreviation “spp.” is usually used to refer to all the individual species within a genus, e.g., Cornusspp. refers to all the plants within the dogwood genus.
species diversity
Almost the same as species richness, but in more technical literature, an ecosystem that is said to be more diverse if the species present have equal population sizes and less diverse if many species are rare and some are very common.
species richness
The number of species within a region. See also: species diversity.
specimen (pl. specimens)

Item considered typical of its class, e.g., an herbarium specimen. 2. Plant grown for landscape exhibition, e.g., a specimen tree.
specimen preparation
The collection, selection, arrangement, pressing, drying, and mounting of a plant for an herbariumspecimen, allowing many specimens to be stored in minimal space.
specimen tree
A tree placed so people can gain the greatest enjoyment for the color, texture, scent, or other pleasures it provides.
spermatium (pl. spermatia)
The nonmotile male cell of red seaweeds; also used for similar cells in fungi.
spermatocyte
A cell which gives rise to sperm cells.
Spermatophyta
A major taxonomic division containing all plants which reproduce by seed, subdivided into Gymnospermae and Angiospermae.
spermatophyte (syn. seed plant)
Any plant of the division Spermatophyta, the higher plants that produce seeds, including the gymnosperms and angiosperms.
spermatozoid
A motile ciliated male reproductive cell.
sphagnum moss
Bog mosses that are often used as soil additives and packing material because of their ability to retain moisture. See also: peat moss.
spherical
Nearly round.
spicate
Arranged in or resembling a spike.
spiciform
Spike-like.
spiculate
Having fine fleshy points.
spider mite (syn. red spider)
Any of several small, red, web-spinning mites of the family Tetranychidae that feed on and damage leaves.
spike
A form of simple inflorescence with the flowers sessile or nearly so upon a more or less elongatedcommon axis.
spike harrows
A field implement with spikes that breaks up clods and levels soil.
spikelet (syn. earlet)
A small or secondary spike, most often refiring to those forming the inflorescence of grasses.
spindle-shaped
Same as fusiform.
spine
A sharp woody or rigid outgrowth from a stem, leaf, or other plant part.
spinescent

Having spines. 2. Terminating in a spine. 3. Modified to form a spine.
spinose
Spine-like, or having spines.
spinulate
Having spinules.
spinule
A minute spine or spine-like process.
spinulose
Thorny; set with small spines.
spired
Having a crown with several points.
splitting maul
An axe with a broad, heavy head, shaped into a wide nonstick wedge.
spoils
Soil from stream or drain bottoms dredged out when the channel was made deeper.
sporangiophore
A specialized stalk bearing a sporangium.
sporangium (pl. sporangia)
A tiny globe in which the spores are produced; often applied to the capsule, but by some authorsrestricted to the spore sac, or inner sac of the capsule containing the spores.
spore
The reproductive organ in cryptogams which in function corresponds to a seed but possesses no embryo.
sporecase
Same as sporangium; the case in which the spores are borne.
sporocarp
The fruit cases of certain cryptogams containing sporangia or spores.
sporogenesis
Reproduction by means of spores.
sporogenous
Describes cells or tissues in which spores are formed.
sporophyll
A leaf bearing spores.
sporophyte (syn. sporophore)

The spore-bearing part or generation. In mosses, it consists of the seta and capsule and constitutes the so-called fruit. In ferns, the conspicuous plants that bear spores.
spororogonium
The sporophyte or spore-bearing part of the moss.
sport
An abrupt, naturally occurring genetic change resulting in a branch that differs in appearance from the rest of the plant, or, a plant derived by propagation from such a genetically changed branch.
spreader

A lawn cart that applies seed, fertilizer, etc. 2. A large wagon used on organic farms to apply manureto the fields.
spreading
Having a horizontally branching habit.
sprig
A small part of a plant, such as stolons used for propagations, twigs bearing flowers, etc.
spring wood
Wood which is formed during the period of rapid growth in the spring, appearing lighter in color than the wood formed later as the growth slows down.
spud (alt. ice spud)
A tool shaped like a flat-ended oar that is used to chop a hole through ice for fishing or studying underwater biology, but which may also be used to clear ice from sidewalks, cut sod or roots, edgesidewalks, etc.
spur
A hollow sac-like or tubular extension of some part of a blossom, usually nectariferous. Also, a short, slow-growing branchlet.
spur branch
A stubby branchlet with densely crowded leaves and leaf scars.
spurred
Calcarate; refers to a flower with tubular projections from the petals or sepals, usually with nectarglands.
sqarulose
Diminutively squarrose.
squamate (alt. squamose, alt. squamous)
Covered with scales.
squamulate
Having squamules.
squamulose
Having small scales.
squarrose
Turned back at a right angle or more sharply, as the tip of a phyllary.
stability
Resistance to variation in populations and disturbances, resulting in the persistence of communitycomposition over long periods of time.
staddle
A foundation of trunk and main branches, either of rootstock or stembuilder, for grafting.
staking
Providing support for newly-planted trees or tall plants by tying the plant to one or more stakes planted beside and parallel to the plant.
stalk
A short, supporting axis.
stalked
Joined to an axis with a stalk.
stamen
One of the male, pollen-bearing organs of the flower.
staminal
Attached to or referring to the stamen.
staminate
With stamens and without pistils.
staminode (syn. staminodium, syn. staminoide)
A sterile stamen, or any structure without anther corresponding to a stamen.
staminoid
A pollenless stamen.
standard (syn. banner)

A plant with a strong vertical stem, particularly one that would not normally grow that way; a vine or shrub trained as a tree with a single trunk and a more or less spherical top. 2. Uppermost, usually erect, petals of some flowers; the upper dilated petal of a papilionaceous corolla.
stegocarpous
Having the capsule operculate.
stele
The axial vascular and associated tissues from the endodermis inward.
stellate (syn. stelliform)
Star-shaped.
stem
The main ascending axis of a plant.
stembuilder
A variety of woody plants used in a graft as intermediate stem piece to provide a strong trunk for standard trees, or to introduce resistance to disease or winter injury.
steppe
A vast arid, usually level and grassland, particularly those of southeastern Europe and Asia and generally having extreme variations in temperature. See also: pampas, prairie.
stereids
The small thick-walled cells seen in.
sterigma (pl. sterigmata)
The spike on which a fungus spore is home.
sterigmata
The small woody stem to which the leaves of spruces and hemlocks are attached.
sterile

Barren. Refers to fern leaves that do not produce spores. 2. Unproductive, as a flower without a pistil, or a stamen without an anther.
stichidium
A small structure (lobe or branch) bearing tetrasporangia.
stigma
That part of a pistil through which fertilization by the pollen is affected.
stigmatic
Belonging to or characteristic of the stigma.
stigmatose
With conspicuous stamens.
stigonema
A filamentous blue-green algae.
stilt-roots
Adventitious support roots.
stimulants
Herbal medicines that can increase the energy or activity of a tissue, organ, etc.
stipel
A stipule of a leaflet.
stipellate
Having stipels.
stipitate
Having a stipe or short stem.
stipular
Belonging to stipules.
stipulate
Having stipules.
stipule
An appendage at the base of a petiole, often appearing in pairs, one on each side, as found on roses.
stipule scar
The scar left on a twig by the fall of a stipule.
stock driveway
A lane; a strip of land designated for the movement of livestock.
stock plants
Mother plants kept for cuttings to reproduce the plant.
stoloniferous
Producing stolons.
stoloniferous stem
A slender creeping stem with minute leaves.
stoloniform
Like a stolon, underground stem.
stoma (pl. stomata)

An orifice in the epidermis of a leaf communicating with internal air cavities. 2. A pore in the wall of a capsule surrounded by special guard-cells and serving the same purpose as the stomata in the epidermis of the leaves of flowering plants.
stone (alt. stone fruit)
The single seed of a drupe, surrounded by a large, hard shell and covered by pulp.
stool
Clump of shoots growing from near the ground.
stooling
Cutting down to ground level to induce tightly packed new growth.
stopping
Pinching off the terminal bud to induce branching.
storm surge
Before and during a storm, an abnormal and rapid rise in lake level along the shore, caused mostly by strong onshore winds and changes of atmospheric pressure.
stove
A heated greenhouse.
straggling
Semiupright.
straight
Describing ray florets having little or no curvature throughout their length (used in describing dahlias.)
strain
A number of plants which have similar characteristics such as color blends, different enough to be designated, but not sufficiently distinct to be considered a horticultural variety.
stramineous
Straw-colored.
strand

The area of lakeshore above the waterline that is subject to the action of wind and rain. 2. On ocean beaches, the area between high and low tidal marks.
stratified
Having prominent horizontal layers.
stratify (n. stratification)
Artificially overcoming a seed’s dormancy by placing them in layers of moisture-retaining media and keeping them under generally cool and moist conditions for a period of time, so as to simulate winter conditions.
strawflowers
Species of plants which easily become dried flowers.
stria (pl. striae)
A thin ridge or groove.
striate (syn. striated)
Marked with fine, longitudinal lines or ridges.
strict
Very straight and upright.
strigose
Beset with appressed sharp straight and stiff hairs.
strigulose
Minutely strigose.
string tags
Waterproof tags used to mark collected specimens for herbarium study.
striolated
With delicate lines or ridges.
strip cropping
A method of planting a crop in parallel bands following the contours of the slope; this lessens erosion. See also: field strip cropping, filter strip.
strobiloid (alt. strobiliform)
Cone-shaped.
strobilus (alt. strobile)

Woody cone, as in Gymnospermae. 2. A cone-shaped inflorescence marked by imbricatedsporophylls. 3. A dense cone-like structure producing spores.
stroma

A compact mass of fungal hyphae producing perithecia or pycnidia. 2. The colorless proteinaceousmatrix of a chloroplast in which the chlorophyll-containing lamellae are embedded.
strone (alt. stron)
A hill that terminates a ridge; the end of a ridge.
strophile
An appendage at the hilum of certain seeds.
strophiole
Resulting from an inverted ovary that has fusion of the nucellus and funiculus. The strophiole is a small swelling on a ridge.
struma
A goiter-like swelling on one side at the base of the capsule.
strumose
Having a struma.
stunt virus
A type of virus disease which stops the growth of a plant.
stylar (alt. styled)
Either having styles or found on a style.
style
The usually attenuated portion of the pistil connecting the stigma and ovary.
stylopodium
A disc-shaped enlargement found at the base of the style in some Umbelliferae.
styptic
A medicine to stop minor surface bleeding, e.g., razor cuts.
sub-
A prefix meaning somewhat or physically below.
subarctic
Of, relating to, or being the regions bordering on the arctic zone.
suberect
Ascending toward the edges.
suberin
A complex fatty substance found especially in the cell walls of cork.
suberization (adj. suberized)
The conversion of the cell walls into corky tissue, called suberin.
suberous
Cork-like.
subgenus
An intermediate category in taxonomy between genus and species.
subimbricate
Somewhat overlapped; slightly shingled.
submergent vegetation
Plants with stems and leaves below water level, sometimes with reproductive parts above water.
submersed (alt. submerged)
Underwater.
subopposite
Almost opposite but one leaf or leaflet of each pair a little above the other.
subsessile
With a slight stalk.
subshrub
A plant with more or less woody branches and roots, less woody than a true shrub, but more woody than a perennial herbaceous plant; a small shrub, woody only at the base.
subspecies (abr. ssp.)
The rank of taxa below species but above variety; a subdivision of a species whose members have certain hereditary characteristics distinct from other populations of that species. A subspecies is added to the specific binomial and preceded by “ssp.”, such as obtusifolia in the epithet Grevillea thelemannianassp. obtusifolia.
substratum (syn. substrate)
The material on which a plant grows.
subtend
To be just below and close up to or enclosed in its axil.
subterranean
Below ground.
subtropical (syn. semitropical)
Of, relating to, or being the regions bordering on the tropical zone.
succession (syn. biotic succession)

The development of a plant community from its initial stage to its climax stage; usually from one consisting of grasses and forbs to one of shrubs and, finally, to forest. However, depending on the climate, the climax stage may consist of a prairie, savana, or any number of other ecosystems. 2. The changes in the species composition of communities following a natural or human disturbance like the natural filling of a pond or the clearing of a road through a forest.
succulent
A plant having fleshy stems or leaves, often adapted to xeric conditions.
sucker
A shoot from the root or lower part of a stem. In roses, a young cane emerging below the bud union and therefore representing the variety of the understock rather than the top variety. A shoot appearing on a tree limb is called a water sprout.
suffrutescent
Slightly or obscurely shrubby.
suffruticose
Very low and woody; diminutively shrubby.
sulcate
Grooved or furrowed.
summer wood
Wood produced late in the growing season when growth is slow.
sunken garden
A garden recessed into the ground to create a secluded atmosphere or to facilitate a view down onto a feature such as a knot garden.
super- (alt. supra-)
A prefix meaning above, greater than, superior to.
superficial
On the surface.
superior ovary
With the flower parts growing from below the ovary.
superparasitism
Parasitization of a host by more than one individual, usually of a single species.
superphosphate
A fertilizer with a very high amount of phosphates, uusally 30-45 percent, and most often used to promote flowering in ornamental plants.
superposed
Accessory buds which occur above the axillary buds.
supersaturation
The uptake of unusual amounts of water by a plant; this can cause plant specimens to mold rather than dry.
supersporangium
A structure containing tetrasporangia and acting as a sporangium.
supra-axillary
Borne above the axil.
suprabasal
Above the base, usually referring to pinnae or pinnules.
supramedial
Beyond the middle.
surculose
Producing suckers.
suspended
Describes an ovule that is hanging from the apex of the cell.
swale
A wet depression between beach ridges, fed by ground water and runoff, rarely influenced by changes in water level.
swell
A huge natural dome structure where movement of the earth’s crust has raised stratigraphic layers in mounds rather than breaking them with a fault. Erosion exposes the oldest layers in the middle with subsequent layers surrounding it in concentric circles.
symbiosis (adj. symbiotic)
The living together of different species of organism which may or may not be to their mutual benefit. See also: mutualism, parasitism.
symmetrical
Describes a flower that can be divided into similar halves.
sympatric
Describes two or more closely-related species having coincident or overlapping ranges of distributionbut not interbreeding. See also: allopatric.
sympodium (pl. sympodia, adj. sympodial)
A primary axis without a single, persistent growing point that develops from a series of lateral branches which change direction in succession and give it a zigzag form, as occurs with orchids of the genusCattleya. See also: monopodium.
syncarp
An aggregate or multiple fruit produced from fused or coherent pistils, the small individual fruitsmassing and growing together into a single fruit, coalescent.
syncarpous
With the carpels of the gynoecium united in a compound ovary.
syncline
A geological structure that appears when strata fold downward toward a common line.
synoicous (alt. synaecious)
Having the male and the female organs mixed together in the same cluster.
synonym (abr. syn.)
Any one of two or more names used for the same taxon rank; a rejected name due to misapplication or difference in taxonomic judgment. See also: basionym, homonym, tautonym, autonym, taxonomic synonym, nomenclatural synonym.
syntype
Any one of two or more specimens cited by the author when no holotype was designated, or when two or more were designated. See also: holotype, lectotype, neotype, isotype, nomenclatural type, topotype.
systematics
The study of the evolutionary and genetic relationships among organisms and of their phenotypesimilarities and differences.
tautonym
An illegitimate binomial in which the genus and species are the same word, such as Amoracia armoracia, later changed to Armoracia rusticana. See also: basionym, homonym, synonym, autonym.
taxon (pl. taxa)
A group of genetically similar organisms that are classified together as a species, genus, family, etc.
taxonomic synonym
Any one of two or more different names based on different plant specimens which were later judged to belong to the same taxon.
taxonomy (adj. taxonomic)
The classification of organisms based on genetic similarities.
temperature summation
The totality of effective temperatures or day-degrees for a specified period of time, especially the time required for an organism to reach maturity, like a summer. See also: aliquote.
temporal

Refers to the sides of the skull behind the eyes, the temples. 2. Refers to time.
tender
Describes a plant easily killed by frost.
tendril
A clasping, twining, slender outgrowth of the stem that helps support climbing plants.
tension
The stress resulting from elongation.
tepal
A division of the perianth–a sepal or a petal–of a flower in which the calyx and corolla are almost identical in appearances, as occurs in tulips and lilies. In Orchidaceae, this term refers to any sepal or petal, except the labellum.
terete
Having a circular transverse section.
terminal
Apical.
ternate
With three nearly equal divisions.
terrace
The entire outdoor paved surface; the term often refers to the patio only.
terrarium
A small artificial environment for a specified habitat. An aquarium is for a water habitat, usually containing fish and plants. A terrarium generally has a woodland setting, with the emphasis on plants rather than animals.
terrestrial
Growing in soil.
tetraploid
Having twice the normal number of chromosomes.
thallus (pl. thalli)
In cryptogams, a cellular expansion taking the place of stem and foliage and forming the main body fungi and lichens.
thatch
A layer of dead grass that does not decay into soil.
thermal stratification
The successive horizontal layers of a body of water having different temperatures, each layer more or less sharply different from the adjoining ones, with the warmest at the top. See also: thermocline, epilimnion, turnover.
thermocline
In bodies of water, there are normally stratified layers of decreasing temperature with increased depth. Thermocline is the layer where the temperature starts to decrease rapidly, about one degree centigrade or more with each meter (3 feet) of depth. See also: epilimnion, thermal stratification.
thorn
A stout, sharp, woody outgrowth of the stem or branch.
throat
The orifice of a gamopetalous corolla or calyx; the part between the proper tube and the limb.
tier
A radial, layered branching habit of excurrent trees.
till
Mixed deposits of gravel, boulders, sand, and other materials that were carried by a glacier.
timber
Standing trees, felled trees, or seasoned logs. See also: lumber, pulpwood.
timber cruising (timber cruiser)
The process of surveying forest land with the objective of predicting how many board feet of lumberthe plot will yield.
tip
The apex.
tip layering
A propagative method similar to air layering, except a flexible branch such as blackberry, Rubus, is bent to the ground and buried in moist soil or a mixture of peat moss and sand, instead of being wrapped on the upright stem.
tissue culture
The growth of small pieces of plant tissue (usually the meristem) under sterile conditions in artificial media.
tooth

The processes comprising the fringe around a peristome. 2. The smallest division of a frond.
toothed
Having the margin shallowly divided into small, tooth-like segments.
top variety
The variety bud-grafted to the understock, and thus the variety that flowers.
topography
The configuration of a surface including its relief and the position of its natural and man-made features.
topotype
A specimen collected later from the original type locality, or from the area from which the type was described. See also: holotype, lectotype, neotype, isotype, nomenclatural type, syntype.
torus
The receptacle of a flower.
trailing
Lying prostrate on the surface or on other vegetation.
translucent
Thin enough to transmit light, but not a concrete image.
transparent
Thin enough to transmit an image.
transpiration
The passage of water through a land plant and usually out the pores of its leaves.
transplant
To move a plant from one place to another.
transverse
Cross-wise in position.
tree
A woody, self-supporting perennial plant usually with a single main stem and generally growing more than 20 feet tall.
tree line (alt. tree limit, timberline)
The height above which trees can no longer survive on a mountainside, or the northern or southern limit beyond which only stunted forms appear on a continent toward a pole.
trench
A long cut, ditch, channel, especially man-made.
tri-
A prefix meaning three.
triangular
Having the outline of a triangle.
trimerous
Having flower parts, such as petals, sepals, and stamens, in groups of three.
tropical
Of, being, or characteristic of a region or climate that is frost-free with temperatures high enough to support year-round plant growth.
true leaf
A leaf typical of a plant that appears subsequent to the cotyledons.
trunk
The major woody stem of a tree.
tuber
A thickened and short subterranean branch having numerous buds or eyes and used for food storage.
tubercle
A small tuber or tuber-like (not necessarily subterranean) body.
tubular
With the petals partly united to form a tube.
tundra
The vegetation type of very cold climates overlying permafrost and consisting of lichens, sedges, mosses, grasses, and low woody plants.
turbinate
Top-shaped; inversely conical.
turnover (syn. overturn, syn. fall overturn)
The cycling of upper and lower strata of water in bodies of fresh water in autumn, and vice versa in spring, caused by the heating or cooling of upper surfaces. When the upper surface is cooler than the lower water, the cool water sinks.
twig
The end subdivision of a branch; a young shoot, generally applied to the growth of the past season.
twining
Clasping by winding around.
twisted
The seta of many mosses twists strongly in drying. If the twist is such as would be made by seizing the capsule and twisting it to the right, it is said to be twisted to the right. It is possible that this twisting of the seta aids in scattering the spores.
type
One of several categories of herbarium specimens, chosen by taxonomists as being close or identical to that chosen by the original author.
ultimate
Of the lowest order; the smallest division.
umbel
An inflorescence in which the peduncles or pedicels of a cluster spring from the same point.
umbellet
A secondary umbel.
underdrainage

The placement of perforated plastic pipe or porous clay pipe underground, often with a layer of crushed stone over it, to carry away excess water. 2. A natural or man-made system allowing excess water to be carried away from a cultivated field.
understock (syn. rhizome, syn. rootstock)
The stock or root plant onto which a shoot has been grafted to produce a new plant in bud-grafting, especially in cases of double-worked trees.
understory
The shrubs and smaller trees between the forest canopy and the ground cover.
undifferentiated
Describes parts or tissues which are very difficult to tell apart, like the tepals of tulips, Tulipa.
unicellular
Made of one cell.
uniform
Always having the same shape, as opposed to dimorphous.
union
A graft.
unisexual
A flower of one sex only, either pistillate (female) or staminate (male).
united
Fused together.
unstratified
Lacking distinct layers.
vacuole
A cavity or vesicle in the cytoplasm of a cell usually containing fluid.
vale
A small valley.
valve
One of the pieces into which a dehiscing capsule splits.
vane
One leaf of a windmill or pinwheel.
variegated
Marked, striped, or blotched with some color in addition to the plant’s general overall color.
variety (alt. varietas, abr. var.)
The rank of taxa below subspecies but above forma; a plant which retains most of the characteristics of the species, but differs in some way such as flower or leaf color, size of mature plant, etc. A variety is added to the specific binomial and preceded by “var.”, such as saxatilis in the epithet Juniperus communisvar. saxatilis.
vascular
Furnished with vessels or ducts.
vegetational analysis
Any of various methods of studying small areas of constituent plants, often counting the numbers of plants of communities, to make extrapolations to a larger area. See also: belt transect, basal area, vegetational cover, timber cruising.
veil
The calyptra.
vein
The thread of fibro-vascular tissue in a leaf or other organ, especially those which branch (as distinguished from nerves).
veinlet
A subdivision or branch of a vein.
veld (alt. veldt)
A term commonly used in South Africa for open country occupied by grasslands in higher elevations and scrub or savanna at lower elevations.
venation (syn. nervation)
The manner in which the veins are arranged in the leaf.
ventral
Belonging to the anterior or inner face of an organ; the opposite of dorsal.
verge
Border, limit, or boundary; a berm.
vernal
Appearing in spring.
vernation
The arrangement of leaves in the bud.
very double
Roses with a great many petals.
vesicle
A small bladder or air-cavity.
vessel
A row of lignified water-conducting cells rendered continuous by the absorption of the crosswalls.
viable
Describes a seed that is capable of germinating.
vicariation
The occurrence of ecological equivalents replacing each other in similar habitats in different geographic areas, e.g., caribou in North America and reindeer in Eurasia.
vine
A plant that trails, clings, or twines, and requires support to grow vertically.
viscous
Not free-flowing; having the consistency of syrup.
vouchering
Furnishing proof, supporting a claim, as of taxonomic study.
Wardian case
A case of glass sides and top used to maintain the humidity necessary for growing tropical and subtropical plants in temperate region buildings. A popular feature of Victorian homes, used often to display ferns.
water sprout
A sucker produced on the trunk, stem, or large branch of a plant, particularly fruit trees, usually growing straight and at a right angle to the axis. These rapidly growing suckers are generally removed to allow a more open tree and to provide better nutrition to the fruit.
watershed
The complete area from which runoff drains to feed a stream or body of water.
wavy
See also: undulate.
weed
A plant growing where it is not wanted, often to the detriment of a crop or the disfigurement of a landscape design.
wetland
An area where saturation or repeated inundation of water is the determining factor in the nature of the soils, as well as the plants and animals living there. Included in the term are marshes, swamps, bogs, fens, bay heads, wet meadows, potholes, sloughs, bayous, river flood plains, estuaries, and lake margins.
whole
Having all its proper parts or components, often describing a flower.
whorl (adj. whorled)
An arrangement of leaves, petals, sepals, etc., in a circle around the stem.
wildfire
A fire that is out of control, such as a forest fire or a burn through a grassland, often threatening houses or farms.
wildflower
A flowering plant growing and usually propagating itself outside of cultivation. Often, but not always, refers to plants native to a region.
windward
The side from which the wind blows. See also: leeward.
wing

A thin, dry, leaf-like membrane found on many fruits, seeds, and leafstalks; any membranous or thin expansion bordering or surrounding an organ. 2. The lateral petal of a papilionaceous corolla.
winged
With projecting thin flat membranes or corky outgrowths.
wood
The hard, fibrous inner tissue of the trunk and branches of a tree or shrub.
woodland (syn. woods)
An area primarily covered with trees.
woody
Forming stems that mature to wood.
woody plant
With the stems and limbs containing lignin.
woolly
Clothed with long and tortuous or matted hairs.
xeric
Of, or adapted to, an extremely dry habitat.
xylem
Tissue specialized for the transport of water and minerals upward through the plant.
zygomorphic
Capable of division by only one plane of symmetry.
zygote
The diploid cell formed by the union of two gametes.